| Class: | angiosperms |
| Order: | Ericales |
| Family: | Pyrolaceae |
| Editor | Rilke 2014 |
| Name acc. to: | APweb |
| Herbar: | list records    |
| Synonym: | Ericaceae (acc. to Heywood 2007) |
| Description: | Evergreen herbs. Leaves coriaceous in radical rosette. Flowers (4-)5-merous; sepals free or united at base; stamens twice as many as petals, obdiplostemon, anthers opening by pores, pollen-grains released in tetrads (except in Orthilia); style 1, stigma entire or (4-)5-lobed. Fruit a (4)5 loculicidal capsule, seeds numerous, small, with two appendages. |
| Tax. Comments: | Often included in expanded Ericaceae due to molecular studies (Heywood 2007). |
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| genus: 3 |
| species: 9 |
| Habit (i)general appearance of a plant | |
| Growth form: (i)Herb, shrub, tree or climber. | herb (i)Herbaceous, erect plant, up to 2m high, mostly with a leafy shoot; if perennial, shoots die to the ground each season, shoots are not woody
example: Artemisia pectinata 
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| Special growth forms or habits: | evergreen (i)Leaves remain on the plant in wintertime or in the dry season
example: Juniperus, Pinus, Ephedra
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| Parasite status: (i)Is the plant a half- or full parasite? | no parasite/saprophyte (i)Plant fully autonomous, leaves with chlorophyll
example: Most plants, Ranunculus
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| Water or terrestrial plant: (i)Where do the plants grow? | terrestrial (i)Plant grows on dry land
example: Orostachys spinosa
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| Leaf (i)expanded, usually photosynthetic organ of a plant (including phylloclades) | |
| Leaf development: (i)Structure and development of leaves. | with green leaves (i)Plant with green leaves
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| Leaf arrangement: (i)Arrangement of leaves at the stem. | basal rosette (i)Leaves positioned at the base of the stem; stem often without leaves, no visible internodes (but flowers often on erect stems, and these may have few leaves)
example: Limonium, Potentilla, Plantago; also used in Liliales with basaly crouwded leaves (Tofieldia, Zigadenus etc.) 
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| Simple or divided leaves: (i)Are the leaves simple or completely divided in several parts? Blade of the leaf entire or (more or less) deeply dissected. Attention: There are various appearances of the leaf margin (from entire to toothed and lobed). Here, we ignore this and ask only for dissections that separate the leaf for more than one third of its length or width, whatever is smaller. Sometimes, it is difficult to tell apart compound leaves from a shoot system with simple leaves: look for stipulae and/or axillary buds at the ground of the leaves: if only some possess these structures, the others are most likely leaflets of a compound leaf. | simple (i)Non-divided leaf, but margin may be incised nearly to the ground 
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| Petiole: (i)Leaf divided into stalk (petiole) and blade. | with (i)Leaves with petiole (stalk) 
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| Stipule: (i)Leaflets at the base of the petiole, these are smaller and of different shape. | none (i)Without stipules
example: Euphorbia, Ericaceae s.l.
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| Leaf colour upper side: (i)Shades of green on the leaf, upper side. | green (i)Clear green
example: Tribulus terrestris
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| Leaf colour lower side: (i)Shades of green on the leaf, lower side. | green (i)Clear green, in most species
example: Angelica decurrens
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| Leaf veination: (i)Arrangement of the main veins of a leaf. | pinnate (i)One main vein, several side veins, sometimes inconspicuous
example: Cicerbita   
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| Flower (i)reproductive portion of the plant, consisting of sepals, petals, stamens, and pistils | |
| Flower appearance and pollination: (i)General appearance of the flower. | attractive, animal-pollinated (i)attractive and coloured flowers, mostly large, attracting surely animals
example: Trollius, Rosa, Chamaerhodos
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| Perianth arrangement: (i)Attention: in some plants, flowers may be dimorphic in different ways (dioecious or gynodioecious). If flowers vary, record the characters of the most showy flowers. | double, different (i)Two types of perianth leaves, differently coloured (sepals: outer periant leaves, usually greenish, and petals: inner perianth leaves, usually coloured)
example: Parnassia  
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| Flower symmetry: (i)Symmetry of the perianth leaves. Attention: to assess this character, look on sepals, petals and stamens, but neglect carpels and ovary. | radiary, regular (actinomorphic) (i)More than two axis of symmetry
example: Saxifraga: 5; Iris: 3 
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| Flower form: (i)common forms of flowers ? Veronica | simple (flat) - Do not confuse with inflorescences as in some Asteraceae (i)Petals spread out, flower appearing flat
example: Mollugo, Trientalis, Pulsatilla, Saxifraga 
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| Sepal number: (i)Number of sepal leaves (outer perianth leaves, calyx leaves, mostly greenish). Attention, this character applies only for flowers separated in sepals and petals, thus excluding most monocots. Be aware of the bracts (involucral leaves) of Asteraceae flowerheads, do not qualify these as sepals! Be also aware in Rosaceae is often an epicalyx developed, in this case count all parts. | 5 (i)
example: Polemonium
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| Sepal fusion: (i)To which degree are the sepal leaves connected? Attention, this character applies only for flowers separated in sepals and petals, thus excluding most monocots. Be aware of the bracts (involucral leaves) of Asteraceae flowerheads, do not qualify these as sepals! | free (i)All leaves separate from each other
example: Geranium
fused at base (i)Leaves with a joint base, but fused over not more than half of the entire length
example: Campanula
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| Petal / Tepal number: (i)Number of petal leaves (inner perianth leaves, usually coloured). | 5 (i)
example: Potentilla
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| Petal / Tepal fusion: (i)To which degree are the petal leaves connected? Petals sympetalous. | free (i)all petal leaves separate from each other
example: Anthriscus
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| Spur: (i)A hollow, slender, sac-like appendage of the perianth leaves, storing nectar. | no spur (i)Flower without appendage
example: Peganum
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| Stamen number: (i)Attention: We ask for the reproductive organs of the flower dispersing pollen. Count only fully fertile stamens, not staminodia (e.g. Parnassia). | 10 (i)
example: Silene
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| Stamen fusion: (i)To which degree are the stamens fused? Attention: Whereas the pollen sacs itself are often free., their stalks (filaments) may be fused. Here, we count them as fused if they are together over at least one thirth of their length. | free (i)Stamens with separate bases
example: Malus
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| Pistil number: (i)Number of pistils (female floral organs: style, if developed; stigma and carpels/ovary together build the pistil). | 1 (i)One carpel, but clearly one stigma
example: Pyrola, Primula, Alyssum
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| Carpel number: (i)Number of carpels (carpel: forming a simple pistil or part of a compound pistil, modified leaf). | 5
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| Carpel fusion: (i)To which degree are the carpels (modified leaf forming simple pistil or part of a compound pistil) fused. | fused (i)Carpels united into an ovary, only styles are free
example: Malus, Berberis
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| Style number: (i)Portion of the pistil connecting the stigma to the ovary. | 1
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| Stigma number per style: (i)Number of stigmas per style. | 1 (i)One stigma, sessile or with a developed style
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| Ovary position: (i)For entirely or partly fused carpels, describe their position in relation to the insertion point of perianth leaves (best done by doing a longitudinal section of a flower). | superior (hypogynous) (i)Base of carpels attached above insertion point of perianth leaves, carpels free or fused
example: Delphinium, Anemone  
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| Sex: (i)Distribution of male and female organs among flowers, only most commonly cases. | bisexual, hermaphrodite (i)All or nearly all flowers of a plant with male and female parts
example: Haplophyllum, Chenopodium
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| Inflorescence (i)flowering part of a plant, describes the arrangement of the flowers on the flowering axis | |
| Inflorescence: (i)Structure of the inflorescence. | Flowers in inflorescence (i)No solitary flowers
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| Inflorescence type: (i)Types of inflorescence. Attention: We here ask for the botanical nomenclature of inflorescences, which is sufficiently complicated. Tick only, if you are certain, or tick all inflorescence types that appear similar of these of the plant in question. | raceme (i)Stalked flowers arranged along a simple main axis, often one by one in the axils of leaves, maturing from bottom upwards
example: Aconitum barbatum 
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| Fruit (i)the seed bearing organ, with or without adnate parts; a ripened ovary and any other structures which are attached and ripen with it. Aggregate fruits are handled like simple fruits for determination. | |
| Consistency: (i)Fleshy fruits or dry fruits, see dispersal adaptations for further classification. | dry (i)With a dry outer shell, no fleshy parts, but seed (embryo) could be edible
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| Type of fruit: (i)Common fruit types (including pseudocarp). | Solitary fruits (i)   
capsule (i)Dry dehiscent fruit, releasing seeds by slits or holes.
example: Poppy, most Caryophyllaceae, Cerastium, a lot of Scrophulariaceae, Iris (oppened capsule looks like Delphinium), Zygophyllum - it is a very common fruit type   
Dehiscent fruits (i)Fruits open along a longitudinale line (except silicula)
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| Size of fruit: (i)Size of the fruit including appendage. | from 5 mm to 10 mm (i)
example: Silene: small capsule opening with teeth
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| Seed number: (i)Estimate the number of seeds per fruit, if recognizable seeds are in the fruit (in rare cases a fruit may contain one seeded nuts: rose hip, carex). | 7-50 (i)Numerable, but may be counted
example: Vaccinum: multi-seeded berry, Ptilotrichum: few-seeded siliqua
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| Root / shoot below ground (i)plant part below ground (in most cases), including below ground shoots, without leaves | |
| Root type: (i)Organisation of the roots. | allorhizous (i)Plant with a conspicuous tap root, one larger tap root with side roots
example: Dicotyledonae  inherited by order Ericales: allorhizous
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