| Class: | angiosperms |
| Order: | Lamiales |
| Family: | Plantaginaceae |
| Editor | S. Rilke, March 2013 |
| Name acc. to: | APGII |
| Herbar: | list records    |
| Description: | Mostly herbaceous plants. Leaves surrounding the base in a basal rosette, or opposite at leafy stems. Inflorescence a compact spike or head. Flowers 4-merous, small, corolla scarious. Fruit a capsule. |
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| genus: 1 |
| species: 8 |
| Habit (i)general appearance of a plant | |
| Growth form: (i)Herb, shrub, tree or climber. | herb (i)Herbaceous, erect plant, up to 2m high, mostly with a leafy shoot; if perennial, shoots die to the ground each season, shoots are not woody
example: Artemisia pectinata 
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| Special growth forms or habits: | rosette plant (i)All or nearly all leaves in a basal rosette, only inflorescences on erect stalks, these without or with much smaller leaves
example: Primula farinosa 
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| Parasite status: (i)Is the plant a half- or full parasite? | no parasite/saprophyte (i)Plant fully autonomous, leaves with chlorophyll
example: Most plants, Ranunculus
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| Water or terrestrial plant: (i)Where do the plants grow? | terrestrial (i)Plant grows on dry land
example: Orostachys spinosa
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| Leaf (i)expanded, usually photosynthetic organ of a plant (including phylloclades) | |
| Leaf arrangement: (i)Arrangement of leaves at the stem. | basal rosette (i)Leaves positioned at the base of the stem; stem often without leaves, no visible internodes (but flowers often on erect stems, and these may have few leaves)
example: Limonium, Potentilla, Plantago; also used in Liliales with basaly crouwded leaves (Tofieldia, Zigadenus etc.) 
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| Simple or divided leaves: (i)Are the leaves simple or completely divided in several parts? Blade of the leaf entire or (more or less) deeply dissected. Attention: There are various appearances of the leaf margin (from entire to toothed and lobed). Here, we ignore this and ask only for dissections that separate the leaf for more than one third of its length or width, whatever is smaller. Sometimes, it is difficult to tell apart compound leaves from a shoot system with simple leaves: look for stipulae and/or axillary buds at the ground of the leaves: if only some possess these structures, the others are most likely leaflets of a compound leaf. | simple (i)Non-divided leaf, but margin may be incised nearly to the ground 
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| Shape of blade: (i)Easy for simple leaves. In compound leaves use the general shape of leaflet. Always check the ground for largest leaves of a plant. To be worked out: how to handle pinnate leaves? | elliptic (including ovate and obovate) (i)Elliptic: broadest at the middle and narrower at the two equal ends; ovate: egg-shaped, attached at the broad end; obovate: attached at the narrower end
example: Limosella aquatica 
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| Leaf apex: (i)Appearance of the tip of leaf resp. leaflets in compound leaves. | obtuse (i)Sides coming together at the apex at an angle greater than 90 degrees
example: Fallopia convolvulus 
acuminate (i)Gradually tapering to a (sharp) point
example: Populus laurifolia? 
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| Leaf margin: (i)Structure of leaf margin (or that of a leaflet in case of compound leaves). Attention: Here we ask for the leaf margin, defined as all those dissections that separate the leaf for less than one third of its length or width, whatever is smaller. To be worked out: how to handle margin of pinnate leaves? | serrate / dentate / crenulate (i)Margin saw-like or rounded teethed
example: Betula, Lophanthus (crenulate) 
finely serrate / dentate (i)Fine teeth, more than 20 per leaf length; usually only one lateral vein per tooth
example: Pedicularis 
entire (i)Plain margin, not toothed
example: Iris 
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| Stipule: (i)Leaflets at the base of the petiole, these are smaller and of different shape. | none (i)Without stipules
example: Euphorbia, Ericaceae s.l.
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| Leaf veination: (i)Arrangement of the main veins of a leaf. | parallel (i)Most veins arranged parallel to the length of leaf, mostly no pronounced main vein (usually in elongate to linear leaves)
example: Most Monocotyledonae, Plantago, Veratrum, a lot of Caryophyllaceae looks like that. 
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| Flower (i)reproductive portion of the plant, consisting of sepals, petals, stamens, and pistils | |
| Flower appearance and pollination: (i)General appearance of the flower. | not attractive, wind-pollinated or some water plants (i)Small, colourless or green flowers
example: Betula, grasslike plants: Carex, Setaria, Juncus
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| Perianth arrangement: (i)Attention: in some plants, flowers may be dimorphic in different ways (dioecious or gynodioecious). If flowers vary, record the characters of the most showy flowers. | double, different (i)Two types of perianth leaves, differently coloured (sepals: outer periant leaves, usually greenish, and petals: inner perianth leaves, usually coloured)
example: Parnassia  
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| Diameter of flower: (i)Diameter of flower or flower head. | to 5 mm (i)
example: Aruncus
from 5 mm to 10 mm (i)
example: Stellaria
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| Flower symmetry: (i)Symmetry of the perianth leaves. Attention: to assess this character, look on sepals, petals and stamens, but neglect carpels and ovary. | radiary, regular (actinomorphic) (i)More than two axis of symmetry
example: Saxifraga: 5; Iris: 3 
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| Flower form: (i)common forms of flowers ? Veronica | tubular to funnel-shaped (i)Petals form a tube, are often partially united to a cylindrical corolla, often surrounded by a calyx 
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| Sepal number: (i)Number of sepal leaves (outer perianth leaves, calyx leaves, mostly greenish). Attention, this character applies only for flowers separated in sepals and petals, thus excluding most monocots. Be aware of the bracts (involucral leaves) of Asteraceae flowerheads, do not qualify these as sepals! Be also aware in Rosaceae is often an epicalyx developed, in this case count all parts. | 4 (i)
example: Sinapis
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| Sepal fusion: (i)To which degree are the sepal leaves connected? Attention, this character applies only for flowers separated in sepals and petals, thus excluding most monocots. Be aware of the bracts (involucral leaves) of Asteraceae flowerheads, do not qualify these as sepals! | free (i)All leaves separate from each other
example: Geranium
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| Petal / Tepal number: (i)Number of petal leaves (inner perianth leaves, usually coloured). | 4 (i)
example: Galium
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| Petal / Tepal fusion: (i)To which degree are the petal leaves connected? Petals sympetalous. | fused at base (i)petal leaves with a joint base, but fused over not more than 50% of the entire length
example: Myosotis, Pedicularis, Cortusa
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| Spur: (i)A hollow, slender, sac-like appendage of the perianth leaves, storing nectar. | no spur (i)Flower without appendage
example: Peganum
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| Stamen number: (i)Attention: We ask for the reproductive organs of the flower dispersing pollen. Count only fully fertile stamens, not staminodia (e.g. Parnassia). | 4 (i)Extremely rare, may be absent
example: Plantago
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| Stamen fusion: (i)To which degree are the stamens fused? Attention: Whereas the pollen sacs itself are often free., their stalks (filaments) may be fused. Here, we count them as fused if they are together over at least one thirth of their length. | free (i)Stamens with separate bases
example: Malus
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| Pistil number: (i)Number of pistils (female floral organs: style, if developed; stigma and carpels/ovary together build the pistil). | 2 (i)Two stigmas, often cleaved like a snakes tongue
example: Salvia, Arnica, Bupleurum, Bromus, Saxifraga, Veronica
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| Carpel fusion: (i)To which degree are the carpels (modified leaf forming simple pistil or part of a compound pistil) fused. | fused (i)Carpels united into an ovary, only styles are free
example: Malus, Berberis
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| Ovary position: (i)For entirely or partly fused carpels, describe their position in relation to the insertion point of perianth leaves (best done by doing a longitudinal section of a flower). | superior (hypogynous) (i)Base of carpels attached above insertion point of perianth leaves, carpels free or fused
example: Delphinium, Anemone  
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| Sex: (i)Distribution of male and female organs among flowers, only most commonly cases. | unisexual (i)
example: Rhodiola
dioecious (i)Male and female flowers at different individuals
example: Antennaria
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| Inflorescence (i)flowering part of a plant, describes the arrangement of the flowers on the flowering axis | |
| Inflorescence: (i)Structure of the inflorescence. | Flowers in inflorescence (i)No solitary flowers
Simple inflorescences (i)Flowers sessile on a main shoot or on short to long not branched side shoots
example: Polygonum bistorta  
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| Appearance: (i)Outer look of the inflorescence. | terminal (i)Inflorescence is the highest point of the plant and may consist of a single flower only
example: Cypripedium, Rhaponticum, Ligularia sibirica, Echinops
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| Inflorescence type: (i)Types of inflorescence. Attention: We here ask for the botanical nomenclature of inflorescences, which is sufficiently complicated. Tick only, if you are certain, or tick all inflorescence types that appear similar of these of the plant in question. | spike (spadix) (i)All flowers sessile and crowded along a main axis, there may be several spikes on a shoot; sometimes axis thickened (spadix)
example: Plantago, Carex vesicaria, Vicia, Typha (spadix)  
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| Fruit (i)the seed bearing organ, with or without adnate parts; a ripened ovary and any other structures which are attached and ripen with it. Aggregate fruits are handled like simple fruits for determination. | |
| Consistency: (i)Fleshy fruits or dry fruits, see dispersal adaptations for further classification. | dry (i)With a dry outer shell, no fleshy parts, but seed (embryo) could be edible
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| Type of fruit: (i)Common fruit types (including pseudocarp). | Solitary fruits (i)   
capsule (i)Dry dehiscent fruit, releasing seeds by slits or holes.
example: Poppy, most Caryophyllaceae, Cerastium, a lot of Scrophulariaceae, Iris (oppened capsule looks like Delphinium), Zygophyllum - it is a very common fruit type   
Dehiscent fruits (i)Fruits open along a longitudinale line (except silicula)
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| Opening of fruit: (i)Mode of dehiscence at maturity to release seeds. | opening / dehiscent (i)Dry? Fruits opening with different types
opening with lid (i)The seed capsule opens with a small cap or cover
example: Plantago, Hyoscyamus, Anagallis (but not in Mongolia) 
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| Size of fruit: (i)Size of the fruit including appendage. | to 5 mm (i)
example: Halerpestes: many folicles forming dry nutlets
from 5 mm to 10 mm (i)
example: Silene: small capsule opening with teeth
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| Seed number: (i)Estimate the number of seeds per fruit, if recognizable seeds are in the fruit (in rare cases a fruit may contain one seeded nuts: rose hip, carex). | 2-6 (i)2-6 single seeds, well recognizable
example: Crataegus: few-seeded berry
7-50 (i)Numerable, but may be counted
example: Vaccinum: multi-seeded berry, Ptilotrichum: few-seeded siliqua
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| Shoot/Stem (i)a young stem or branch | |
| Spines, thorns or prickles: (i)Shoot with conspicuous spines, thorns or prickles. | absent (i)Stem glabrous or hairy, but never with spines, thornes or prickles
example: Gentiana barbata
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| Root / shoot below ground (i)plant part below ground (in most cases), including below ground shoots, without leaves | |
| Root type: (i)Organisation of the roots. | allorhizous (i)Plant with a conspicuous tap root, one larger tap root with side roots
example: Dicotyledonae  inherited by order Lamiales: allorhizous
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