| Class: | angiosperms |
| Order: | Lamiales |
| Family: | Plantaginaceae |
| Genus: | Plantago |
| Herbar: | list records    |
| Description: | Leaves in radical rosette, simple mostly entire. Flowers small, tetramerous, in dense spicate inflorescence. |
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| species: 8 |
| Habit (i)general appearance of a plant | |
| Growth form: (i)Herb, shrub, tree or climber. | herb (i)Herbaceous, erect plant, up to 2m high, mostly with a leafy shoot; if perennial, shoots die to the ground each season, shoots are not woody
example: Artemisia pectinata   inherited by family Plantaginaceae: herb
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| Special growth forms or habits: | rosette plant (i)All or nearly all leaves in a basal rosette, only inflorescences on erect stalks, these without or with much smaller leaves
example: Primula farinosa  inherited by family Plantaginaceae: rosette plant
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| Parasite status: (i)Is the plant a half- or full parasite? | no parasite/saprophyte (i)Plant fully autonomous, leaves with chlorophyll
example: Most plants, Ranunculus inherited by family Plantaginaceae: no parasite/saprophyte
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| Water or terrestrial plant: (i)Where do the plants grow? | terrestrial (i)Plant grows on dry land
example: Orostachys spinosa inherited by family Plantaginaceae: terrestrial
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| Leaf (i)expanded, usually photosynthetic organ of a plant (including phylloclades) | |
| Leaf arrangement: (i)Arrangement of leaves at the stem. | basal rosette (i)Leaves positioned at the base of the stem; stem often without leaves, no visible internodes (but flowers often on erect stems, and these may have few leaves)
example: Limonium, Potentilla, Plantago; also used in Liliales with basaly crouwded leaves (Tofieldia, Zigadenus etc.)  inherited by family Plantaginaceae: basal rosette
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| Simple or divided leaves: (i)Are the leaves simple or completely divided in several parts? Blade of the leaf entire or (more or less) deeply dissected. Attention: There are various appearances of the leaf margin (from entire to toothed and lobed). Here, we ignore this and ask only for dissections that separate the leaf for more than one third of its length or width, whatever is smaller. Sometimes, it is difficult to tell apart compound leaves from a shoot system with simple leaves: look for stipulae and/or axillary buds at the ground of the leaves: if only some possess these structures, the others are most likely leaflets of a compound leaf. | simple (i)Non-divided leaf, but margin may be incised nearly to the ground  inherited by family Plantaginaceae: simple
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| Shape of blade: (i)Easy for simple leaves. In compound leaves use the general shape of leaflet. Always check the ground for largest leaves of a plant. To be worked out: how to handle pinnate leaves? | elliptic (including ovate and obovate) (i)Elliptic: broadest at the middle and narrower at the two equal ends; ovate: egg-shaped, attached at the broad end; obovate: attached at the narrower end
example: Limosella aquatica  inherited by family Plantaginaceae: elliptic (including ovate and obovate)
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| Leaf apex: (i)Appearance of the tip of leaf resp. leaflets in compound leaves. | obtuse (i)Sides coming together at the apex at an angle greater than 90 degrees
example: Fallopia convolvulus  inherited by family Plantaginaceae: obtuse
acuminate (i)Gradually tapering to a (sharp) point
example: Populus laurifolia?  inherited by family Plantaginaceae: acuminate
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| Leaf margin: (i)Structure of leaf margin (or that of a leaflet in case of compound leaves). Attention: Here we ask for the leaf margin, defined as all those dissections that separate the leaf for less than one third of its length or width, whatever is smaller. To be worked out: how to handle margin of pinnate leaves? | serrate / dentate / crenulate (i)Margin saw-like or rounded teethed
example: Betula, Lophanthus (crenulate)  inherited by family Plantaginaceae: serrate / dentate / crenulate
finely serrate / dentate (i)Fine teeth, more than 20 per leaf length; usually only one lateral vein per tooth
example: Pedicularis  inherited by family Plantaginaceae: finely serrate / dentate
entire (i)Plain margin, not toothed
example: Iris  inherited by family Plantaginaceae: entire
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| Stipule: (i)Leaflets at the base of the petiole, these are smaller and of different shape. | none (i)Without stipules
example: Euphorbia, Ericaceae s.l. inherited by family Plantaginaceae: none
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| Leaf veination: (i)Arrangement of the main veins of a leaf. | parallel (i)Most veins arranged parallel to the length of leaf, mostly no pronounced main vein (usually in elongate to linear leaves)
example: Most Monocotyledonae, Plantago, Veratrum, a lot of Caryophyllaceae looks like that.  inherited by family Plantaginaceae: parallel
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| Flower (i)reproductive portion of the plant, consisting of sepals, petals, stamens, and pistils | |
| Flower appearance and pollination: (i)General appearance of the flower. | not attractive, wind-pollinated or some water plants (i)Small, colourless or green flowers
example: Betula, grasslike plants: Carex, Setaria, Juncus inherited by family Plantaginaceae: not attractive, wind-pollinated or some water plants
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| Perianth arrangement: (i)Attention: in some plants, flowers may be dimorphic in different ways (dioecious or gynodioecious). If flowers vary, record the characters of the most showy flowers. | double, different (i)Two types of perianth leaves, differently coloured (sepals: outer periant leaves, usually greenish, and petals: inner perianth leaves, usually coloured)
example: Parnassia   inherited by family Plantaginaceae: double, different
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| Diameter of flower: (i)Diameter of flower or flower head. | to 5 mm (i)
example: Aruncus inherited by family Plantaginaceae:
from 5 mm to 10 mm (i)
example: Stellaria inherited by family Plantaginaceae:
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| Flower symmetry: (i)Symmetry of the perianth leaves. Attention: to assess this character, look on sepals, petals and stamens, but neglect carpels and ovary. | radiary, regular (actinomorphic) (i)More than two axis of symmetry
example: Saxifraga: 5; Iris: 3  inherited by family Plantaginaceae: radiary, regular (actinomorphic)
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| Flower form: (i)common forms of flowers ? Veronica | tubular to funnel-shaped (i)Petals form a tube, are often partially united to a cylindrical corolla, often surrounded by a calyx  inherited by family Plantaginaceae: tubular to funnel-shaped
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| Sepal number: (i)Number of sepal leaves (outer perianth leaves, calyx leaves, mostly greenish). Attention, this character applies only for flowers separated in sepals and petals, thus excluding most monocots. Be aware of the bracts (involucral leaves) of Asteraceae flowerheads, do not qualify these as sepals! Be also aware in Rosaceae is often an epicalyx developed, in this case count all parts. | 4 (i)
example: Sinapis inherited by family Plantaginaceae: 4
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| Sepal fusion: (i)To which degree are the sepal leaves connected? Attention, this character applies only for flowers separated in sepals and petals, thus excluding most monocots. Be aware of the bracts (involucral leaves) of Asteraceae flowerheads, do not qualify these as sepals! | free (i)All leaves separate from each other
example: Geranium inherited by family Plantaginaceae: free
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| Petal / Tepal number: (i)Number of petal leaves (inner perianth leaves, usually coloured). | 4 (i)
example: Galium inherited by family Plantaginaceae: 4
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| Petal / Tepal fusion: (i)To which degree are the petal leaves connected? Petals sympetalous. | fused at base (i)petal leaves with a joint base, but fused over not more than 50% of the entire length
example: Myosotis, Pedicularis, Cortusa inherited by family Plantaginaceae: fused at base
fused (i)petal leaves united, only tips are free (gamopetalous, sympetalous)
example: Linnaea, Adenophora, Stellera
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| Spur: (i)A hollow, slender, sac-like appendage of the perianth leaves, storing nectar. | no spur (i)Flower without appendage
example: Peganum inherited by family Plantaginaceae: no spur
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| Stamen number: (i)Attention: We ask for the reproductive organs of the flower dispersing pollen. Count only fully fertile stamens, not staminodia (e.g. Parnassia). | 4 (i)Extremely rare, may be absent
example: Plantago inherited by family Plantaginaceae: 4
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| Stamen fusion: (i)To which degree are the stamens fused? Attention: Whereas the pollen sacs itself are often free., their stalks (filaments) may be fused. Here, we count them as fused if they are together over at least one thirth of their length. | free (i)Stamens with separate bases
example: Malus inherited by family Plantaginaceae: free
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| Pistil number: (i)Number of pistils (female floral organs: style, if developed; stigma and carpels/ovary together build the pistil). | 2 (i)Two stigmas, often cleaved like a snakes tongue
example: Salvia, Arnica, Bupleurum, Bromus, Saxifraga, Veronica inherited by family Plantaginaceae: 2
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| Carpel fusion: (i)To which degree are the carpels (modified leaf forming simple pistil or part of a compound pistil) fused. | fused (i)Carpels united into an ovary, only styles are free
example: Malus, Berberis inherited by family Plantaginaceae: fused
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| Ovary position: (i)For entirely or partly fused carpels, describe their position in relation to the insertion point of perianth leaves (best done by doing a longitudinal section of a flower). | superior (hypogynous) (i)Base of carpels attached above insertion point of perianth leaves, carpels free or fused
example: Delphinium, Anemone   inherited by family Plantaginaceae: superior (hypogynous)
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| Sex: (i)Distribution of male and female organs among flowers, only most commonly cases. | bisexual, hermaphrodite (i)All or nearly all flowers of a plant with male and female parts
example: Haplophyllum, Chenopodium
unisexual (i)
example: Rhodiola inherited by family Plantaginaceae: unisexual
dioecious (i)Male and female flowers at different individuals
example: Antennaria inherited by family Plantaginaceae: dioecious
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| Inflorescence (i)flowering part of a plant, describes the arrangement of the flowers on the flowering axis | |
| Inflorescence: (i)Structure of the inflorescence. | Flowers in inflorescence (i)No solitary flowers inherited by family Plantaginaceae: Flowers in inflorescence
Simple inflorescences (i)Flowers sessile on a main shoot or on short to long not branched side shoots
example: Polygonum bistorta   inherited by family Plantaginaceae: Simple inflorescences
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| Appearance: (i)Outer look of the inflorescence. | terminal (i)Inflorescence is the highest point of the plant and may consist of a single flower only
example: Cypripedium, Rhaponticum, Ligularia sibirica, Echinops inherited by family Plantaginaceae: terminal
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| Inflorescence type: (i)Types of inflorescence. Attention: We here ask for the botanical nomenclature of inflorescences, which is sufficiently complicated. Tick only, if you are certain, or tick all inflorescence types that appear similar of these of the plant in question. | spike (spadix) (i)All flowers sessile and crowded along a main axis, there may be several spikes on a shoot; sometimes axis thickened (spadix)
example: Plantago, Carex vesicaria, Vicia, Typha (spadix)   inherited by family Plantaginaceae: spike (spadix)
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| Fruit (i)the seed bearing organ, with or without adnate parts; a ripened ovary and any other structures which are attached and ripen with it. Aggregate fruits are handled like simple fruits for determination. | |
| Consistency: (i)Fleshy fruits or dry fruits, see dispersal adaptations for further classification. | dry (i)With a dry outer shell, no fleshy parts, but seed (embryo) could be edible inherited by family Plantaginaceae: dry
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| Type of fruit: (i)Common fruit types (including pseudocarp). | Solitary fruits (i)    inherited by family Plantaginaceae: Solitary fruits
capsule (i)Dry dehiscent fruit, releasing seeds by slits or holes.
example: Poppy, most Caryophyllaceae, Cerastium, a lot of Scrophulariaceae, Iris (oppened capsule looks like Delphinium), Zygophyllum - it is a very common fruit type    inherited by family Plantaginaceae: capsule
Dehiscent fruits (i)Fruits open along a longitudinale line (except silicula) inherited by family Plantaginaceae: Dehiscent fruits
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| Opening of fruit: (i)Mode of dehiscence at maturity to release seeds. | opening / dehiscent (i)Dry? Fruits opening with different types inherited by family Plantaginaceae: opening / dehiscent
opening with lid (i)The seed capsule opens with a small cap or cover
example: Plantago, Hyoscyamus, Anagallis (but not in Mongolia)  inherited by family Plantaginaceae: opening with lid
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| Size of fruit: (i)Size of the fruit including appendage. | to 5 mm (i)
example: Halerpestes: many folicles forming dry nutlets inherited by family Plantaginaceae:
from 5 mm to 10 mm (i)
example: Silene: small capsule opening with teeth inherited by family Plantaginaceae:
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| Dispersal: (i)Appearance of fruit or seed (if single) and adaptations to dispersal. | Otherwise (i)All parts dry, no conspicuous adaptations
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| Seed number: (i)Estimate the number of seeds per fruit, if recognizable seeds are in the fruit (in rare cases a fruit may contain one seeded nuts: rose hip, carex). | 2-6 (i)2-6 single seeds, well recognizable
example: Crataegus: few-seeded berry inherited by family Plantaginaceae: 2-6
7-50 (i)Numerable, but may be counted
example: Vaccinum: multi-seeded berry, Ptilotrichum: few-seeded siliqua inherited by family Plantaginaceae: 7-50
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| Shoot/Stem (i)a young stem or branch | |
| Spines, thorns or prickles: (i)Shoot with conspicuous spines, thorns or prickles. | absent (i)Stem glabrous or hairy, but never with spines, thornes or prickles
example: Gentiana barbata inherited by family Plantaginaceae: absent
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| Root / shoot below ground (i)plant part below ground (in most cases), including below ground shoots, without leaves | |
| Root type: (i)Organisation of the roots. | allorhizous (i)Plant with a conspicuous tap root, one larger tap root with side roots
example: Dicotyledonae inherited by order Lamiales: allorhizous
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