| Class: | angiosperms |
| Order: | Rosales |
| Family: | Rosaceae |
| Genus: | Crataegus |
| Herbar: | list records  |
| Description: | Trees and shrubs 2-4 m high, with thorns (C. dahurica without thorns). Leaves lobed. Inflorescences many-flowered, corymbose. Fruits apple-like with 3-4 stones, red. Differentiation on herbarium sheets is very difficult, clear cut features are missing. |
| Tax. Comments: | Due to multiple hybridization a reliable determination is not possible in all cases. Especially differentiation on herbarium sheets is very difficult, since clear cut features are missing. For determination, leaves from short shoots should only be used. |
| Link to Flora of China: | http://www.efloras.org/florataxon.aspx?flora_id=2&taxon_id=108272 |
| open map in a new window |  |
| species: 3 |
| Habit (i)general appearance of a plant | |
| Growth form: (i)Herb, shrub, tree or climber. | herb (i)Herbaceous, erect plant, up to 2m high, mostly with a leafy shoot; if perennial, shoots die to the ground each season, shoots are not woody
example: Artemisia pectinata 
perennial (i)Living for several to many years, as opposed to annual and biennial
shrub, subshrub or semishrub (i)Shrub, multi-stemmed, mostly (0.2) 0.5 - 5 m high, shoots woody up to the tip
example: Caragana leucophloea 
tree (i)Woody plant with a clear main trunk, at least 2-3 m tall
example: Ulmus pumila 
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| Size of plant: (i)Attention: use flowering or fruiting specimens to assess plant height (many biennial plants possess only a basal rosette in the first year). | from 1000 mm to 3000 mm
more than 3000 mm
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| Parasite status: (i)Is the plant a half- or full parasite? | no parasite/saprophyte (i)Plant fully autonomous, leaves with chlorophyll
example: Most plants, Ranunculus  inherited by family Rosaceae: no parasite/saprophyte
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| Water or terrestrial plant: (i)Where do the plants grow? | terrestrial (i)Plant grows on dry land
example: Orostachys spinosa
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| Leaf (i)expanded, usually photosynthetic organ of a plant (including phylloclades) | |
| Leaf development: (i)Structure and development of leaves. | with green leaves (i)Plant with green leaves inherited by family Rosaceae: with green leaves
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| Leaf arrangement: (i)Arrangement of leaves at the stem. | alternate (i)One leaf per node; distiche: arranged in two vertical rows, equitant
example: Phragmites  
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| Simple or divided leaves: (i)Are the leaves simple or completely divided in several parts? Blade of the leaf entire or (more or less) deeply dissected. Attention: There are various appearances of the leaf margin (from entire to toothed and lobed). Here, we ignore this and ask only for dissections that separate the leaf for more than one third of its length or width, whatever is smaller. Sometimes, it is difficult to tell apart compound leaves from a shoot system with simple leaves: look for stipulae and/or axillary buds at the ground of the leaves: if only some possess these structures, the others are most likely leaflets of a compound leaf. | simple (i)Non-divided leaf, but margin may be incised nearly to the ground 
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| Leaf margin: (i)Structure of leaf margin (or that of a leaflet in case of compound leaves). Attention: Here we ask for the leaf margin, defined as all those dissections that separate the leaf for less than one third of its length or width, whatever is smaller. To be worked out: how to handle margin of pinnate leaves? | serrate / dentate / crenulate (i)Margin saw-like or rounded teethed
example: Betula, Lophanthus (crenulate) 
lobed (i)Curved division, be aware of deeply lobed leaves than compare with palmatisected (character: leaf division)
example: Crataegus, Dryas, Rubus humulifolius 
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| Petiole: (i)Leaf divided into stalk (petiole) and blade. | with (i)Leaves with petiole (stalk) 
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| Stipule: (i)Leaflets at the base of the petiole, these are smaller and of different shape. | pair (i)A pair of free stipulae
example: Lathyrus, Trifolium 
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| Leaf colour upper side: (i)Shades of green on the leaf, upper side. | green (i)Clear green
example: Tribulus terrestris
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| Leaf colour lower side: (i)Shades of green on the leaf, lower side. | green (i)Clear green, in most species
example: Angelica decurrens
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| Leaf veination: (i)Arrangement of the main veins of a leaf. | pinnate (i)One main vein, several side veins, sometimes inconspicuous
example: Cicerbita   
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| Flower (i)reproductive portion of the plant, consisting of sepals, petals, stamens, and pistils | |
| Flower appearance and pollination: (i)General appearance of the flower. | attractive, animal-pollinated (i)attractive and coloured flowers, mostly large, attracting surely animals
example: Trollius, Rosa, Chamaerhodos inherited by family Rosaceae: attractive, animal-pollinated
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| Perianth arrangement: (i)Attention: in some plants, flowers may be dimorphic in different ways (dioecious or gynodioecious). If flowers vary, record the characters of the most showy flowers. | double, different (i)Two types of perianth leaves, differently coloured (sepals: outer periant leaves, usually greenish, and petals: inner perianth leaves, usually coloured)
example: Parnassia  
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| Flower symmetry: (i)Symmetry of the perianth leaves. Attention: to assess this character, look on sepals, petals and stamens, but neglect carpels and ovary. | radiary, regular (actinomorphic) (i)More than two axis of symmetry
example: Saxifraga: 5; Iris: 3  inherited by family Rosaceae: radiary, regular (actinomorphic)
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| Flower form: (i)common forms of flowers ? Veronica | simple (flat) - Do not confuse with inflorescences as in some Asteraceae (i)Petals spread out, flower appearing flat
example: Mollugo, Trientalis, Pulsatilla, Saxifraga  inherited by family Rosaceae: simple (flat) - Do not confuse with inflorescences as in some Asteraceae
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| Sepal number: (i)Number of sepal leaves (outer perianth leaves, calyx leaves, mostly greenish). Attention, this character applies only for flowers separated in sepals and petals, thus excluding most monocots. Be aware of the bracts (involucral leaves) of Asteraceae flowerheads, do not qualify these as sepals! Be also aware in Rosaceae is often an epicalyx developed, in this case count all parts. | 5 (i)
example: Polemonium
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| Petal / Tepal number: (i)Number of petal leaves (inner perianth leaves, usually coloured). | 5 (i)
example: Potentilla
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| Petal / Tepal fusion: (i)To which degree are the petal leaves connected? Petals sympetalous. | free (i)all petal leaves separate from each other
example: Anthriscus inherited by family Rosaceae: free
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| Spur: (i)A hollow, slender, sac-like appendage of the perianth leaves, storing nectar. | no spur (i)Flower without appendage
example: Peganum inherited by family Rosaceae: no spur
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| Stamen number: (i)Attention: We ask for the reproductive organs of the flower dispersing pollen. Count only fully fertile stamens, not staminodia (e.g. Parnassia). | > 10 (i)
example: Nymphaea, Callianthemum, Rosa
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| Stamen fusion: (i)To which degree are the stamens fused? Attention: Whereas the pollen sacs itself are often free., their stalks (filaments) may be fused. Here, we count them as fused if they are together over at least one thirth of their length. | free (i)Stamens with separate bases
example: Malus inherited by family Rosaceae: free
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| Carpel number: (i)Number of carpels (carpel: forming a simple pistil or part of a compound pistil, modified leaf). | 5
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| Ovary position: (i)For entirely or partly fused carpels, describe their position in relation to the insertion point of perianth leaves (best done by doing a longitudinal section of a flower). | inferior (i)Ovary below the point where perianth leaves are inserted, always fused to an ovary
example: Vaccinum  
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| Sex: (i)Distribution of male and female organs among flowers, only most commonly cases. | bisexual, hermaphrodite (i)All or nearly all flowers of a plant with male and female parts
example: Haplophyllum, Chenopodium
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| Inflorescence (i)flowering part of a plant, describes the arrangement of the flowers on the flowering axis | |
| Inflorescence: (i)Structure of the inflorescence. | Flowers in inflorescence (i)No solitary flowers
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| Appearance: (i)Outer look of the inflorescence. | terminal (i)Inflorescence is the highest point of the plant and may consist of a single flower only
example: Cypripedium, Rhaponticum, Ligularia sibirica, Echinops
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| Inflorescence type: (i)Types of inflorescence. Attention: We here ask for the botanical nomenclature of inflorescences, which is sufficiently complicated. Tick only, if you are certain, or tick all inflorescence types that appear similar of these of the plant in question. | others (in traits_comments nicht aufgeführt) (i)Not as above
example: Sparganium: globose capitate
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| Fruit (i)the seed bearing organ, with or without adnate parts; a ripened ovary and any other structures which are attached and ripen with it. Aggregate fruits are handled like simple fruits for determination. | |
| Consistency: (i)Fleshy fruits or dry fruits, see dispersal adaptations for further classification. | fleshy (i)Usually non dehiscent; with a fleshy (swollen because of a high water content) outer shell, flesh edible at least for animals
example: Prunus, Amygdalus, Malus
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| Type of fruit: (i)Common fruit types (including pseudocarp). | pomelike fruit (i)Some (often about 5) fruitlets (follicles) surrounded by fleshy receptacle
example: Malus, Crataegus, Cotoneaster   
Aggregated fruits (i)   
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| Dispersal: (i)Appearance of fruit or seed (if single) and adaptations to dispersal. | Fleshy, edible (i)At least parts with a soft tissue edible at least for some animals (not necessarily for humans)
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| Seed number: (i)Estimate the number of seeds per fruit, if recognizable seeds are in the fruit (in rare cases a fruit may contain one seeded nuts: rose hip, carex). | 2-6 (i)2-6 single seeds, well recognizable
example: Crataegus: few-seeded berry
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| Shoot/Stem (i)a young stem or branch | |
| Spines, thorns or prickles: (i)Shoot with conspicuous spines, thorns or prickles. | spines or thornes (i)Sharp pointed woody structure originating from the plant (thornes derived from a reduced branch, spines from leaves)
example: Prunus spinosa no in Mongolia
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| Root / shoot below ground (i)plant part below ground (in most cases), including below ground shoots, without leaves | |
| Root type: (i)Organisation of the roots. | allorhizous (i)Plant with a conspicuous tap root, one larger tap root with side roots
example: Dicotyledonae inherited by order Rosales: allorhizous
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