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Class:angiosperms
Order:Apiales
Family:Apiaceae
Genus:Angelica
Scientific name: Angelica sylvestris L.
Name acc. to:Gubanov 1996
Herbar:list records    photo available    
Description:Leaflets "sitting" without petiol.
Link to Flora of China:http://www.efloras.org/browse.aspx?flora_id=2&name_str=Angelica+sylvestris
open map in a new windowtaxon distribution for Angelica sylvestris acc. to Geobotanical Regions of Mongolia by Grubov (1955)
Habitat:In forests, meadows (Plants of Central Asia, Vol. 10, 2005).
Habit (i)general appearance of a plant
Growth form: (i)Herb, shrub, tree or climber.herb (i)Herbaceous, erect plant, up to 2m high, mostly with a leafy shoot; if perennial, shoots die to the ground each season, shoots are not woody
example: Artemisia pectinata
inherited by family Apiaceae: herb
Parasite status: (i)Is the plant a half- or full parasite?no parasite/saprophyte (i)Plant fully autonomous, leaves with chlorophyll
example: Most plants, Ranunculus
inherited by family Apiaceae: no parasite/saprophyte
Water or terrestrial plant: (i)Where do the plants grow?terrestrial (i)Plant grows on dry land
example: Orostachys spinosa
inherited by family Apiaceae: terrestrial
Leaf (i)expanded, usually photosynthetic organ of a plant (including phylloclades)
Leaf development: (i)Structure and development of leaves.flattened blade (i)Cross-section of lamina flat, plain inherited by genus Angelica: flattened blade
common leaf (i)Green, often divided in blade and petiole
example: Cotoneaster
inherited by genus Angelica: common leaf
with green leaves (i)Plant with green leaves inherited by family Apiaceae: with green leaves
Leaf arrangement: (i)Arrangement of leaves at the stem.alternate (i)One leaf per node; distiche: arranged in two vertical rows, equitant
example: Phragmites
inherited by family Apiaceae: alternate
Simple or divided leaves: (i)Are the leaves simple or completely divided in several parts? Blade of the leaf entire or (more or less) deeply dissected. Attention: There are various appearances of the leaf margin (from entire to toothed and lobed). Here, we ignore this and ask only for dissections that separate the leaf for more than one third of its length or width, whatever is smaller. Sometimes, it is difficult to tell apart compound leaves from a shoot system with simple leaves: look for stipulae and/or axillary buds at the ground of the leaves: if only some possess these structures, the others are most likely leaflets of a compound leaf.simple (i)Non-divided leaf, but margin may be incised nearly to the ground inherited by family Apiaceae: simple
trifoliate (i)Three-parted leaf (often called ternate)
example: Trifolium, Thermopsis
inherited by family Apiaceae: trifoliate
compound (i)Composed of several similar parts inherited by family Apiaceae: compound inherited by genus Angelica: compound
pinnate (i)With leaflets on both sides of an axis, at least 2 pairs or more
example: Onobrychis
inherited by family Apiaceae: pinnate inherited by genus Angelica: pinnate
bipinnate (i)Twofold pinnate (two levels of pinnae) inherited by family Apiaceae: bipinnate inherited by genus Angelica: bipinnate
complex divided (i)Divided into leaflets of more than second order
example: Artemisia gemelinii, Pulsatilla bungeana
inherited by genus Angelica: complex divided
Petiole: (i)Leaf divided into stalk (petiole) and blade.with (i)Leaves with petiole (stalk) inherited by family Apiaceae: with
Stipule: (i)Leaflets at the base of the petiole, these are smaller and of different shape.none (i)Without stipules
example: Euphorbia, Ericaceae s.l.
inherited by family Apiaceae: none
Leaf colour upper side: (i)Shades of green on the leaf, upper side.green (i)Clear green
example: Tribulus terrestris
inherited by family Apiaceae: green
Leaf colour lower side: (i)Shades of green on the leaf, lower side.green (i)Clear green, in most species
example: Angelica decurrens
inherited by genus Angelica: green
Leaf veination: (i)Arrangement of the main veins of a leaf.pinnate (i)One main vein, several side veins, sometimes inconspicuous
example: Cicerbita
inherited by genus Angelica: pinnate
Flower (i)reproductive portion of the plant, consisting of sepals, petals, stamens, and pistils
Flower appearance and pollination: (i)General appearance of the flower.attractive, animal-pollinated (i)attractive and coloured flowers, mostly large, attracting surely animals
example: Trollius, Rosa, Chamaerhodos
inherited by family Apiaceae: attractive, animal-pollinated
Flower colour: (i)Attention: assess colour of the most colourful parts of the flower, but not of the stamens; be aware of single plants with a mutation (mostly white) on flower colour.greenish (i)petals absent or not distinctly different from colours of leaves, only stigmas (white) or anthers (yellow) may differ in color
example: Chenopodium, Triglochin

white (i)Most plants of the population white
example: Pleurospermum, Maianthemum
inherited by family Apiaceae: white
yellow to orange (i)Pale to golden yellow
example: Ranunculus, Crepis
inherited by genus Angelica: yellow to orange
pink (i)Between red and white
example: Centaurium

Perianth arrangement: (i)Attention: in some plants, flowers may be dimorphic in different ways (dioecious or gynodioecious). If flowers vary, record the characters of the most showy flowers.simple, similar (i)Only one type of perianth leaves (tepals)
example: Tulipa
inherited by family Apiaceae: simple, similar
double, different (i)Two types of perianth leaves, differently coloured (sepals: outer periant leaves, usually greenish, and petals: inner perianth leaves, usually coloured)
example: Parnassia
inherited by family Apiaceae: double, different
Diameter of flower: (i)Diameter of flower or flower head. from 5 mm to 10 mm (i)
example: Stellaria
inherited by family Apiaceae:
Flower symmetry: (i)Symmetry of the perianth leaves. Attention: to assess this character, look on sepals, petals and stamens, but neglect carpels and ovary.radiary, regular (actinomorphic) (i)More than two axis of symmetry
example: Saxifraga: 5; Iris: 3
inherited by family Apiaceae: radiary, regular (actinomorphic)
Flower form: (i)common forms of flowers ? Veronicasimple (flat) - Do not confuse with inflorescences as in some Asteraceae (i)Petals spread out, flower appearing flat
example: Mollugo, Trientalis, Pulsatilla, Saxifraga
inherited by family Apiaceae: simple (flat) - Do not confuse with inflorescences as in some Asteraceae
Sepal number: (i)Number of sepal leaves (outer perianth leaves, calyx leaves, mostly greenish). Attention, this character applies only for flowers separated in sepals and petals, thus excluding most monocots. Be aware of the bracts (involucral leaves) of Asteraceae flowerheads, do not qualify these as sepals! Be also aware in Rosaceae is often an epicalyx developed, in this case count all parts.none or rudimentary (i)Hardly visible or absent, since perianth uniform
example: All monocots with uniform perianth, many Asteraceae and Apiaceae
inherited by family Apiaceae: none or rudimentary inherited by genus Angelica: none or rudimentary
5 (i)
example: Polemonium
inherited by family Apiaceae: 5
Sepal fusion: (i)To which degree are the sepal leaves connected? Attention, this character applies only for flowers separated in sepals and petals, thus excluding most monocots. Be aware of the bracts (involucral leaves) of Asteraceae flowerheads, do not qualify these as sepals!free (i)All leaves separate from each other
example: Geranium
inherited by family Apiaceae: free
Petal / Tepal number: (i)Number of petal leaves (inner perianth leaves, usually coloured).5 (i)
example: Potentilla
inherited by family Apiaceae: 5
Petal / Tepal fusion: (i)To which degree are the petal leaves connected? Petals sympetalous.free (i)all petal leaves separate from each other
example: Anthriscus
inherited by family Apiaceae: free
Spur: (i)A hollow, slender, sac-like appendage of the perianth leaves, storing nectar.no spur (i)Flower without appendage
example: Peganum
inherited by family Apiaceae: no spur
Stamen number: (i)Attention: We ask for the reproductive organs of the flower dispersing pollen. Count only fully fertile stamens, not staminodia (e.g. Parnassia).5 (i)
example: Peucedanum
inherited by family Apiaceae: 5
Stamen fusion: (i)To which degree are the stamens fused? Attention: Whereas the pollen sacs itself are often free., their stalks (filaments) may be fused. Here, we count them as fused if they are together over at least one thirth of their length.free (i)Stamens with separate bases
example: Malus
inherited by family Apiaceae: free
Pistil number: (i)Number of pistils (female floral organs: style, if developed; stigma and carpels/ovary together build the pistil).1 (i)One carpel, but clearly one stigma
example: Pyrola, Primula, Alyssum
inherited by family Apiaceae: 1
Carpel number: (i)Number of carpels (carpel: forming a simple pistil or part of a compound pistil, modified leaf).2 inherited by family Apiaceae: 2
Carpel fusion: (i)To which degree are the carpels (modified leaf forming simple pistil or part of a compound pistil) fused.fused (i)Carpels united into an ovary, only styles are free
example: Malus, Berberis
inherited by family Apiaceae: fused
Style number: (i)Portion of the pistil connecting the stigma to the ovary.2 inherited by family Apiaceae: 2
Stigma number per style: (i)Number of stigmas per style.2 (i)Two stigmas, resulting from two fused carpels with or without developed style inherited by family Apiaceae: 2
Ovary position: (i)For entirely or partly fused carpels, describe their position in relation to the insertion point of perianth leaves (best done by doing a longitudinal section of a flower).inferior (i)Ovary below the point where perianth leaves are inserted, always fused to an ovary
example: Vaccinum
inherited by family Apiaceae: inferior
Sex: (i)Distribution of male and female organs among flowers, only most commonly cases.bisexual, hermaphrodite (i)All or nearly all flowers of a plant with male and female parts
example: Haplophyllum, Chenopodium
inherited by family Apiaceae: bisexual, hermaphrodite
unisexual (i)
example: Rhodiola
inherited by family Apiaceae: unisexual
monoecious (i)Male and female flowers at the same plant
example: Xanthium, Larix, Atriplex
inherited by family Apiaceae: monoecious
Inflorescence (i)flowering part of a plant, describes the arrangement of the flowers on the flowering axis
Inflorescence: (i)Structure of the inflorescence.Flowers in inflorescence (i)No solitary flowers inherited by family Apiaceae: Flowers in inflorescence
Appearance: (i)Outer look of the inflorescence.terminal (i)Inflorescence is the highest point of the plant and may consist of a single flower only
example: Cypripedium, Rhaponticum, Ligularia sibirica, Echinops
inherited by family Apiaceae: terminal
Inflorescence type: (i)Types of inflorescence. Attention: We here ask for the botanical nomenclature of inflorescences, which is sufficiently complicated. Tick only, if you are certain, or tick all inflorescence types that appear similar of these of the plant in question.umbel (i)Flowers on short to long, leafless stalks emerging from one point
example: Allium
inherited by genus Angelica: umbel
double umbel (i)As an umbel, but additional with second order umbels
example: Apiaceae
inherited by genus Angelica: double umbel
Fruit (i)the seed bearing organ, with or without adnate parts; a ripened ovary and any other structures which are attached and ripen with it. Aggregate fruits are handled like simple fruits for determination.
Consistency: (i)Fleshy fruits or dry fruits, see dispersal adaptations for further classification.dry (i)With a dry outer shell, no fleshy parts, but seed (embryo) could be edible inherited by family Apiaceae: dry
Type of fruit: (i)Common fruit types (including pseudocarp).Indehiscent fruits inherited by family Apiaceae: Indehiscent fruits
achene (i)A small, dry, indehiscent fruit with a single seed
example: Asteraceae, Apiaceae (schizocarp), Dipsacaceae, Rosaceae, Ranunculaceae
inherited by family Apiaceae: achene
nut or nutlet (i)Dry fruit with a single, hard stone inside (and usually a large often edible embryo) inherited by family Apiaceae: nut or nutlet
Opening of fruit: (i)Mode of dehiscence at maturity to release seeds.not opening / indehiscent (i)Fruits remain closed at maturity and disperse with seeds inside
example: Corylus (nut), Vaccinium (berry)
inherited by family Apiaceae: not opening / indehiscent
Splitting of fruit: (i)Is a solitary fruit splitting into fruitlets? The fruit splits into separate one-seeded segments at maturity (schizocarp).splitting inherited by family Apiaceae: splitting
in 2 parts (i)
example: Apiaceae
inherited by family Apiaceae: in 2 parts
Size of fruit: (i)Size of the fruit including appendage. to 5 mm (i)
example: Halerpestes: many folicles forming dry nutlets
inherited by family Apiaceae:
from 5 mm to 10 mm (i)
example: Silene: small capsule opening with teeth
inherited by family Apiaceae:
Seed number: (i)Estimate the number of seeds per fruit, if recognizable seeds are in the fruit (in rare cases a fruit may contain one seeded nuts: rose hip, carex).2 (i)Exactly two seeds (or nutlets)
example: Apiaceae
inherited by family Apiaceae: 2
Shoot/Stem (i)a young stem or branch
Spines, thorns or prickles: (i)Shoot with conspicuous spines, thorns or prickles.absent (i)Stem glabrous or hairy, but never with spines, thornes or prickles
example: Gentiana barbata

Root / shoot below ground (i)plant part below ground (in most cases), including below ground shoots, without leaves
Root type: (i)Organisation of the roots.allorhizous (i)Plant with a conspicuous tap root, one larger tap root with side roots
example: Dicotyledonae
inherited by order Apiales: allorhizous
Runners: (i)Plant must be excavated; shoots, subterranean shoots connected by runners.none (i)No runners or stolos visible
example: Dactylorhiza
inherited by family Apiaceae: none
Storage in below-ground structures: (i)Rhizomes or bulbs.beet-like thickened taproot (i)The upper part of the root is clearly thicker than the shoot base
example: Scorzonera
inherited by family Apiaceae: beet-like thickened taproot
bulbs with roots on lower side (i)Leaves at the base of the plant form a bulb, containing one to many leaves
example: Lilium, Allium, Gagea
inherited by family Apiaceae: bulbs with roots on lower side
Distribution (i)region where the plant is likely to be found
Distribution (Veg. Zones): (i)acc. to Grubov 1952Khobdo (i)In distribution data often named as '6'
acc. to: Gubanov 1996
Plant Use
Medical Plant Use: (i)Only if plant is used as medical plantParts used: herb,root and seeds (i)Parts of Plants used in medicine
Ingredients: the herb contains flavonoids (quercetin,diosmin), the leaves yield rutin and quercetin, the plant yields acids, angelic acid, resin,tannin,pectin, coumarins, the root and fruits contain coumarins (i)Medically effective ingredients
Use: The herb is used in Mongolian medicine as an essential constituent of remedies used for treatment of amenorrhea, fall and blow wounds, swelling ulcers,edemas and dropsy. In Chinese medicine, the plant is used to treat fevers and in disposition due to influenza,cephalalgia,water depletion,rheumatism,carbunculosis and abscess. In Russian folk medicine, the root and seeds are used to treat scorbut, depression of activity of digestive organs, anuria, bronchial catarrh, blennothorax. The essential oil, tincture and decoction obtained from this plant are applied in Europe as an orexigenic and spasmolytic to cure hyporexia, flatulence,dysmenorrheal and gastrointestinal apasm. (i)Medical use in eastern and western medicine
acc. to: Ligaa et al. 2009
General Use:medical plant