| Class: | angiosperms |
| Order: | Rosales |
| Family: | Rosaceae |
| Genus: | Padus |
| Scientific name: | Padus avium Mill. |
| Name acc. to: | Gubanov 1996 |
| Herbar: | list records  |
| Synonym: | P. asiatica Kom. (acc. to Grubov 1982) |
| Description: | Shrub or tree up to 10 m tall, annual shoots velvety-pubescent. Leaves elliptical, short-acuminate, beneath pubescent or with rusty hairs in vein axils, later glabrous. Racemes 10-15 cm long, flowers white,highly fragrant. Fruits black, 7-8 mm in diameter. |
| Tax. Comments: | In Flora of China online var. avium, var. pubescens and var. asiatica are differentiated. |
| Link to Flora of China: | http://www.efloras.org/browse.aspx?flora_id=2&name_str=Padus+avium |
| open map in a new window |  |
| Habitat: | River valleys, uremas, river banks in birch kolki, bottom and slopes of creek valleys (Grubov 2001). |
| Habit (i)general appearance of a plant | |
| Growth form: (i)Herb, shrub, tree or climber. | shrub, subshrub or semishrub (i)Shrub, multi-stemmed, mostly (0.2) 0.5 - 5 m high, shoots woody up to the tip
example: Caragana leucophloea   inherited by genus Padus: shrub, subshrub or semishrub
tree (i)Woody plant with a clear main trunk, at least 2-3 m tall
example: Ulmus pumila  inherited by genus Padus: tree
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| Smell & Touch: (i)General appearance of the plant. | odor (i)Plant with an obvious scent
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| Size of plant: (i)Attention: use flowering or fruiting specimens to assess plant height (many biennial plants possess only a basal rosette in the first year). | from 1000 mm to 3000 mm
more than 3000 mm
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| Parasite status: (i)Is the plant a half- or full parasite? | no parasite/saprophyte (i)Plant fully autonomous, leaves with chlorophyll
example: Most plants, Ranunculus inherited by family Rosaceae: no parasite/saprophyte
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| Water or terrestrial plant: (i)Where do the plants grow? | terrestrial (i)Plant grows on dry land
example: Orostachys spinosa
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| Leaf (i)expanded, usually photosynthetic organ of a plant (including phylloclades) | |
| Leaf development: (i)Structure and development of leaves. | with green leaves (i)Plant with green leaves inherited by family Rosaceae: with green leaves
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| Leaf arrangement: (i)Arrangement of leaves at the stem. | alternate (i)One leaf per node; distiche: arranged in two vertical rows, equitant
example: Phragmites  
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| Simple or divided leaves: (i)Are the leaves simple or completely divided in several parts? Blade of the leaf entire or (more or less) deeply dissected. Attention: There are various appearances of the leaf margin (from entire to toothed and lobed). Here, we ignore this and ask only for dissections that separate the leaf for more than one third of its length or width, whatever is smaller. Sometimes, it is difficult to tell apart compound leaves from a shoot system with simple leaves: look for stipulae and/or axillary buds at the ground of the leaves: if only some possess these structures, the others are most likely leaflets of a compound leaf. | simple (i)Non-divided leaf, but margin may be incised nearly to the ground 
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| Shape of blade: (i)Easy for simple leaves. In compound leaves use the general shape of leaflet. Always check the ground for largest leaves of a plant. To be worked out: how to handle pinnate leaves? | elliptic (including ovate and obovate) (i)Elliptic: broadest at the middle and narrower at the two equal ends; ovate: egg-shaped, attached at the broad end; obovate: attached at the narrower end
example: Limosella aquatica 
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| Length of leaves: (i)How long is the leaf, be carefull in compound leaves, measure the complete leaf. | more than 50 mm
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| Leaf apex: (i)Appearance of the tip of leaf resp. leaflets in compound leaves. | acuminate (i)Gradually tapering to a (sharp) point
example: Populus laurifolia? 
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| Leaf margin: (i)Structure of leaf margin (or that of a leaflet in case of compound leaves). Attention: Here we ask for the leaf margin, defined as all those dissections that separate the leaf for less than one third of its length or width, whatever is smaller. To be worked out: how to handle margin of pinnate leaves? | serrate / dentate / crenulate (i)Margin saw-like or rounded teethed
example: Betula, Lophanthus (crenulate) 
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| Leaf base: (i)The angle the leaf blade forms with a real or imaginary (sessile leaves) petiole. Attention: Here, we consider only the base of leaves or leaflets (in case of compound leaves). | cordate (i)Heart shaped, with lobes 
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| Petiole: (i)Leaf divided into stalk (petiole) and blade. | with (i)Leaves with petiole (stalk) 
shorter than blade (i)Petiol shorter than leaf blade
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| Stipule: (i)Leaflets at the base of the petiole, these are smaller and of different shape. | pair (i)A pair of free stipulae
example: Lathyrus, Trifolium  inherited by genus Padus: pair
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| Leaf veination: (i)Arrangement of the main veins of a leaf. | pinnate (i)One main vein, several side veins, sometimes inconspicuous
example: Cicerbita   
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| Flower (i)reproductive portion of the plant, consisting of sepals, petals, stamens, and pistils | |
| Flower appearance and pollination: (i)General appearance of the flower. | attractive, animal-pollinated (i)attractive and coloured flowers, mostly large, attracting surely animals
example: Trollius, Rosa, Chamaerhodos inherited by family Rosaceae: attractive, animal-pollinated
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| Flower colour: (i)Attention: assess colour of the most colourful parts of the flower, but not of the stamens; be aware of single plants with a mutation (mostly white) on flower colour. | white (i)Most plants of the population white
example: Pleurospermum, Maianthemum
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| Perianth arrangement: (i)Attention: in some plants, flowers may be dimorphic in different ways (dioecious or gynodioecious). If flowers vary, record the characters of the most showy flowers. | double, different (i)Two types of perianth leaves, differently coloured (sepals: outer periant leaves, usually greenish, and petals: inner perianth leaves, usually coloured)
example: Parnassia   inherited by genus Padus: double, different
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| Diameter of flower: (i)Diameter of flower or flower head. | to 5 mm (i)
example: Aruncus
from 5 mm to 10 mm (i)
example: Stellaria
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| Flower symmetry: (i)Symmetry of the perianth leaves. Attention: to assess this character, look on sepals, petals and stamens, but neglect carpels and ovary. | radiary, regular (actinomorphic) (i)More than two axis of symmetry
example: Saxifraga: 5; Iris: 3  inherited by family Rosaceae: radiary, regular (actinomorphic)
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| Flower form: (i)common forms of flowers ? Veronica | simple (flat) - Do not confuse with inflorescences as in some Asteraceae (i)Petals spread out, flower appearing flat
example: Mollugo, Trientalis, Pulsatilla, Saxifraga  inherited by family Rosaceae: simple (flat) - Do not confuse with inflorescences as in some Asteraceae
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| Sepal number: (i)Number of sepal leaves (outer perianth leaves, calyx leaves, mostly greenish). Attention, this character applies only for flowers separated in sepals and petals, thus excluding most monocots. Be aware of the bracts (involucral leaves) of Asteraceae flowerheads, do not qualify these as sepals! Be also aware in Rosaceae is often an epicalyx developed, in this case count all parts. | 5 (i)
example: Polemonium inherited by genus Padus: 5
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| Petal / Tepal number: (i)Number of petal leaves (inner perianth leaves, usually coloured). | 5 (i)
example: Potentilla inherited by genus Padus: 5
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| Petal / Tepal fusion: (i)To which degree are the petal leaves connected? Petals sympetalous. | free (i)all petal leaves separate from each other
example: Anthriscus inherited by family Rosaceae: free inherited by genus Padus: free
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| Spur: (i)A hollow, slender, sac-like appendage of the perianth leaves, storing nectar. | no spur (i)Flower without appendage
example: Peganum inherited by family Rosaceae: no spur
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| Stamen number: (i)Attention: We ask for the reproductive organs of the flower dispersing pollen. Count only fully fertile stamens, not staminodia (e.g. Parnassia). | > 10 (i)
example: Nymphaea, Callianthemum, Rosa
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| Stamen fusion: (i)To which degree are the stamens fused? Attention: Whereas the pollen sacs itself are often free., their stalks (filaments) may be fused. Here, we count them as fused if they are together over at least one thirth of their length. | free (i)Stamens with separate bases
example: Malus inherited by family Rosaceae: free
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| Pistil number: (i)Number of pistils (female floral organs: style, if developed; stigma and carpels/ovary together build the pistil). | 1 (i)One carpel, but clearly one stigma
example: Pyrola, Primula, Alyssum
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| Style number: (i)Portion of the pistil connecting the stigma to the ovary. | 1
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| Stigma number per style: (i)Number of stigmas per style. | 1 (i)One stigma, sessile or with a developed style
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| Sex: (i)Distribution of male and female organs among flowers, only most commonly cases. | bisexual, hermaphrodite (i)All or nearly all flowers of a plant with male and female parts
example: Haplophyllum, Chenopodium
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| Inflorescence (i)flowering part of a plant, describes the arrangement of the flowers on the flowering axis | |
| Inflorescence: (i)Structure of the inflorescence. | Flowers in inflorescence (i)No solitary flowers
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| Appearance: (i)Outer look of the inflorescence. | axillary (i)Usually several inflorescences in axillary shoots or single flowers in leaf axils, main shoot remains mostly leafy
example: Tragopogon, Aconogonon
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| Inflorescence type: (i)Types of inflorescence. Attention: We here ask for the botanical nomenclature of inflorescences, which is sufficiently complicated. Tick only, if you are certain, or tick all inflorescence types that appear similar of these of the plant in question. | raceme (i)Stalked flowers arranged along a simple main axis, often one by one in the axils of leaves, maturing from bottom upwards
example: Aconitum barbatum 
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| Fruit (i)the seed bearing organ, with or without adnate parts; a ripened ovary and any other structures which are attached and ripen with it. Aggregate fruits are handled like simple fruits for determination. | |
| Consistency: (i)Fleshy fruits or dry fruits, see dispersal adaptations for further classification. | fleshy (i)Usually non dehiscent; with a fleshy (swollen because of a high water content) outer shell, flesh edible at least for animals
example: Prunus, Amygdalus, Malus inherited by genus Padus: fleshy
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| Type of fruit: (i)Common fruit types (including pseudocarp). | Indehiscent fruits inherited by genus Padus: Indehiscent fruits
Solitary fruits (i)    inherited by genus Padus: Solitary fruits
drupe (stone fruit) (i)Fleshy, indehiscent fruit with a single, hard stone inside
example: Plum, cherry  inherited by genus Padus: drupe (stone fruit)
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| Opening of fruit: (i)Mode of dehiscence at maturity to release seeds. | not opening / indehiscent (i)Fruits remain closed at maturity and disperse with seeds inside
example: Corylus (nut), Vaccinium (berry)
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| Size of fruit: (i)Size of the fruit including appendage. | from 5 mm to 10 mm (i)
example: Silene: small capsule opening with teeth
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| Dispersal: (i)Appearance of fruit or seed (if single) and adaptations to dispersal. | Fleshy, edible (i)At least parts with a soft tissue edible at least for some animals (not necessarily for humans) inherited by genus Padus: Fleshy, edible
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| Seed number: (i)Estimate the number of seeds per fruit, if recognizable seeds are in the fruit (in rare cases a fruit may contain one seeded nuts: rose hip, carex). | 1 (i)A single seed (stone) or seed and fruit wall tightly connected
example: Prunus, Amygdalus: drupe
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| Root / shoot below ground (i)plant part below ground (in most cases), including below ground shoots, without leaves | |
| Root type: (i)Organisation of the roots. | allorhizous (i)Plant with a conspicuous tap root, one larger tap root with side roots
example: Dicotyledonae inherited by order Rosales: allorhizous
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| Distribution (i)region where the plant is likely to be found | |
| Distribution (Veg. Zones): (i)acc. to Grubov 1952 | Khentei (i)In distribution data often named as '2' 
Khangai (i)In distribution data often named as '3' 
Mongol-Daurian (i)In distribution data often named as '4' 
Great Khingan (i)In distribution data often named as '5' 
East Mongolia (i)In distribution data often named as '9' 
acc. to: Gubanov 1996 |
| Distribution Khangay: (i)acc. Flora Khangaya 1989 | I
III
IV
V
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