| Class: | angiosperms |
| Order: | Rosales |
| Family: | Rosaceae |
| Genus: | Rubus |
| Scientific name: | Rubus humulifolius C. A. Mey. |
| Name acc. to: | Gubanov 1996 |
| Herbar: | list records |
| Tax. Comments: | FloraGREIF accessible material of Mongolia (HAL, GFW) does not include this species. |
| Link to Flora of China: | http://www.efloras.org/browse.aspx?flora_id=2&name_str=Rubus+humulifolius |
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| Habitat: | Coniferous and mixed forests and their fringes (Grubov 2001). |
| Habit (i)general appearance of a plant | |
| Growth form: (i)Herb, shrub, tree or climber. | herb (i)Herbaceous, erect plant, up to 2m high, mostly with a leafy shoot; if perennial, shoots die to the ground each season, shoots are not woody
example: Artemisia pectinata   inherited by genus Rubus: herb
perennial (i)Living for several to many years, as opposed to annual and biennial inherited by genus Rubus: perennial
shrub, subshrub or semishrub (i)Shrub, multi-stemmed, mostly (0.2) 0.5 - 5 m high, shoots woody up to the tip
example: Caragana leucophloea  inherited by genus Rubus: shrub, subshrub or semishrub
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| Parasite status: (i)Is the plant a half- or full parasite? | no parasite/saprophyte (i)Plant fully autonomous, leaves with chlorophyll
example: Most plants, Ranunculus inherited by family Rosaceae: no parasite/saprophyte
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| Water or terrestrial plant: (i)Where do the plants grow? | terrestrial (i)Plant grows on dry land
example: Orostachys spinosa inherited by genus Rubus: terrestrial
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| Leaf (i)expanded, usually photosynthetic organ of a plant (including phylloclades) | |
| Leaf development: (i)Structure and development of leaves. | with green leaves (i)Plant with green leaves inherited by family Rosaceae: with green leaves
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| Leaf arrangement: (i)Arrangement of leaves at the stem. | alternate (i)One leaf per node; distiche: arranged in two vertical rows, equitant
example: Phragmites   inherited by genus Rubus: alternate
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| Simple or divided leaves: (i)Are the leaves simple or completely divided in several parts? Blade of the leaf entire or (more or less) deeply dissected. Attention: There are various appearances of the leaf margin (from entire to toothed and lobed). Here, we ignore this and ask only for dissections that separate the leaf for more than one third of its length or width, whatever is smaller. Sometimes, it is difficult to tell apart compound leaves from a shoot system with simple leaves: look for stipulae and/or axillary buds at the ground of the leaves: if only some possess these structures, the others are most likely leaflets of a compound leaf. | simple (i)Non-divided leaf, but margin may be incised nearly to the ground  inherited by genus Rubus: simple
trifoliate (i)Three-parted leaf (often called ternate)
example: Trifolium, Thermopsis  inherited by genus Rubus: trifoliate
compound (i)Composed of several similar parts inherited by genus Rubus: compound
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| Leaf apex: (i)Appearance of the tip of leaf resp. leaflets in compound leaves. | acuminate (i)Gradually tapering to a (sharp) point
example: Populus laurifolia?  inherited by genus Rubus: acuminate
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| Leaf margin: (i)Structure of leaf margin (or that of a leaflet in case of compound leaves). Attention: Here we ask for the leaf margin, defined as all those dissections that separate the leaf for less than one third of its length or width, whatever is smaller. To be worked out: how to handle margin of pinnate leaves? | lobed (i)Curved division, be aware of deeply lobed leaves than compare with palmatisected (character: leaf division)
example: Crataegus, Dryas, Rubus humulifolius  inherited by genus Rubus: lobed
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| Petiole: (i)Leaf divided into stalk (petiole) and blade. | with (i)Leaves with petiole (stalk)  inherited by genus Rubus: with
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| Stipule: (i)Leaflets at the base of the petiole, these are smaller and of different shape. | pair (i)A pair of free stipulae
example: Lathyrus, Trifolium  inherited by genus Rubus: pair
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| Leaf colour lower side: (i)Shades of green on the leaf, lower side. | green (i)Clear green, in most species
example: Angelica decurrens inherited by genus Rubus: green
whitish (i)Nearly white or cream, often (due to dense pubescence)
example: Rubus canescens inherited by genus Rubus: whitish
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| Leaf veination: (i)Arrangement of the main veins of a leaf. | pinnate (i)One main vein, several side veins, sometimes inconspicuous
example: Cicerbita    inherited by genus Rubus: pinnate
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| Flower (i)reproductive portion of the plant, consisting of sepals, petals, stamens, and pistils | |
| Flower appearance and pollination: (i)General appearance of the flower. | attractive, animal-pollinated (i)attractive and coloured flowers, mostly large, attracting surely animals
example: Trollius, Rosa, Chamaerhodos inherited by family Rosaceae: attractive, animal-pollinated
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| Flower colour: (i)Attention: assess colour of the most colourful parts of the flower, but not of the stamens; be aware of single plants with a mutation (mostly white) on flower colour. | white (i)Most plants of the population white
example: Pleurospermum, Maianthemum inherited by genus Rubus: white
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| Perianth arrangement: (i)Attention: in some plants, flowers may be dimorphic in different ways (dioecious or gynodioecious). If flowers vary, record the characters of the most showy flowers. | double, different (i)Two types of perianth leaves, differently coloured (sepals: outer periant leaves, usually greenish, and petals: inner perianth leaves, usually coloured)
example: Parnassia   inherited by genus Rubus: double, different
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| Flower symmetry: (i)Symmetry of the perianth leaves. Attention: to assess this character, look on sepals, petals and stamens, but neglect carpels and ovary. | radiary, regular (actinomorphic) (i)More than two axis of symmetry
example: Saxifraga: 5; Iris: 3  inherited by family Rosaceae: radiary, regular (actinomorphic)
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| Flower form: (i)common forms of flowers ? Veronica | simple (flat) - Do not confuse with inflorescences as in some Asteraceae (i)Petals spread out, flower appearing flat
example: Mollugo, Trientalis, Pulsatilla, Saxifraga  inherited by family Rosaceae: simple (flat) - Do not confuse with inflorescences as in some Asteraceae
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| Sepal number: (i)Number of sepal leaves (outer perianth leaves, calyx leaves, mostly greenish). Attention, this character applies only for flowers separated in sepals and petals, thus excluding most monocots. Be aware of the bracts (involucral leaves) of Asteraceae flowerheads, do not qualify these as sepals! Be also aware in Rosaceae is often an epicalyx developed, in this case count all parts. | 5 (i)
example: Polemonium inherited by genus Rubus: 5
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| Petal / Tepal number: (i)Number of petal leaves (inner perianth leaves, usually coloured). | 5 (i)
example: Potentilla inherited by genus Rubus: 5
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| Petal / Tepal fusion: (i)To which degree are the petal leaves connected? Petals sympetalous. | free (i)all petal leaves separate from each other
example: Anthriscus inherited by family Rosaceae: free
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| Spur: (i)A hollow, slender, sac-like appendage of the perianth leaves, storing nectar. | no spur (i)Flower without appendage
example: Peganum inherited by family Rosaceae: no spur
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| Stamen number: (i)Attention: We ask for the reproductive organs of the flower dispersing pollen. Count only fully fertile stamens, not staminodia (e.g. Parnassia). | > 10 (i)
example: Nymphaea, Callianthemum, Rosa inherited by genus Rubus: > 10
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| Stamen fusion: (i)To which degree are the stamens fused? Attention: Whereas the pollen sacs itself are often free., their stalks (filaments) may be fused. Here, we count them as fused if they are together over at least one thirth of their length. | free (i)Stamens with separate bases
example: Malus inherited by family Rosaceae: free
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| Carpel number: (i)Number of carpels (carpel: forming a simple pistil or part of a compound pistil, modified leaf). | > 5 inherited by genus Rubus: > 5
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| Ovary position: (i)For entirely or partly fused carpels, describe their position in relation to the insertion point of perianth leaves (best done by doing a longitudinal section of a flower). | superior (hypogynous) (i)Base of carpels attached above insertion point of perianth leaves, carpels free or fused
example: Delphinium, Anemone   inherited by genus Rubus: superior (hypogynous)
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| Sex: (i)Distribution of male and female organs among flowers, only most commonly cases. | bisexual, hermaphrodite (i)All or nearly all flowers of a plant with male and female parts
example: Haplophyllum, Chenopodium inherited by genus Rubus: bisexual, hermaphrodite
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| Inflorescence (i)flowering part of a plant, describes the arrangement of the flowers on the flowering axis | |
| Inflorescence: (i)Structure of the inflorescence. | Flowers in inflorescence (i)No solitary flowers inherited by genus Rubus: Flowers in inflorescence
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| Appearance: (i)Outer look of the inflorescence. | terminal (i)Inflorescence is the highest point of the plant and may consist of a single flower only
example: Cypripedium, Rhaponticum, Ligularia sibirica, Echinops inherited by genus Rubus: terminal
axillary (i)Usually several inflorescences in axillary shoots or single flowers in leaf axils, main shoot remains mostly leafy
example: Tragopogon, Aconogonon inherited by genus Rubus: axillary
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| Inflorescence type: (i)Types of inflorescence. Attention: We here ask for the botanical nomenclature of inflorescences, which is sufficiently complicated. Tick only, if you are certain, or tick all inflorescence types that appear similar of these of the plant in question. | others (in traits_comments nicht aufgeführt) (i)Not as above
example: Sparganium: globose capitate inherited by genus Rubus: others (in traits_comments nicht aufgeführt)
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| Fruit (i)the seed bearing organ, with or without adnate parts; a ripened ovary and any other structures which are attached and ripen with it. Aggregate fruits are handled like simple fruits for determination. | |
| Consistency: (i)Fleshy fruits or dry fruits, see dispersal adaptations for further classification. | fleshy (i)Usually non dehiscent; with a fleshy (swollen because of a high water content) outer shell, flesh edible at least for animals
example: Prunus, Amygdalus, Malus inherited by genus Rubus: fleshy
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| Type of fruit: (i)Common fruit types (including pseudocarp). | aggregated drupes, nuts or berries (i)Aggregation of solitary fruits for complex spreading units
example: Fragaria, Rubus  inherited by genus Rubus: aggregated drupes, nuts or berries
Aggregated fruits (i)    inherited by genus Rubus: Aggregated fruits
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| Opening of fruit: (i)Mode of dehiscence at maturity to release seeds. | not opening / indehiscent (i)Fruits remain closed at maturity and disperse with seeds inside
example: Corylus (nut), Vaccinium (berry) inherited by genus Rubus: not opening / indehiscent
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| Size of fruit: (i)Size of the fruit including appendage. | from 5 mm to 10 mm (i)
example: Silene: small capsule opening with teeth inherited by genus Rubus:
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| Dispersal: (i)Appearance of fruit or seed (if single) and adaptations to dispersal. | Fleshy, edible (i)At least parts with a soft tissue edible at least for some animals (not necessarily for humans) inherited by genus Rubus: Fleshy, edible
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| Hairs | |
| Has hairs?: | has hairs inherited by genus Rubus: has hairs
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| Hairs: (i)Appearance, structure, coverage of hairs on plant. | on leaf (i)Hairs on upper side, lower side or on margin of leaf inherited by genus Rubus: on leaf
appearance: soft (i)Hairs very flexible and soft, lay down at a touch
example: Rubus inherited by genus Rubus: appearance: soft
leaf lower side inherited by genus Rubus: leaf lower side
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| Shoot/Stem (i)a young stem or branch | |
| Spines, thorns or prickles: (i)Shoot with conspicuous spines, thorns or prickles. | absent (i)Stem glabrous or hairy, but never with spines, thornes or prickles
example: Gentiana barbata inherited by genus Rubus: absent
spines or thornes (i)Sharp pointed woody structure originating from the plant (thornes derived from a reduced branch, spines from leaves)
example: Prunus spinosa no in Mongolia inherited by genus Rubus: spines or thornes
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| Root / shoot below ground (i)plant part below ground (in most cases), including below ground shoots, without leaves | |
| Root type: (i)Organisation of the roots. | allorhizous (i)Plant with a conspicuous tap root, one larger tap root with side roots
example: Dicotyledonae inherited by order Rosales: allorhizous
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| Runners: (i)Plant must be excavated; shoots, subterranean shoots connected by runners. | none (i)No runners or stolos visible
example: Dactylorhiza inherited by genus Rubus: none
long root suckers or rhizomes (i)Plants grow new shoots from roots or subterranean shoots, called rhizomes, these spacing stems apart
example: Hippophae, Artemisia sericea inherited by genus Rubus: long root suckers or rhizomes
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| Distribution (i)region where the plant is likely to be found | |
| Distribution (Veg. Zones): (i)acc. to Grubov 1952 | Khubsgul (i)In distribution data often named as '1' 
Khentei (i)In distribution data often named as '2' 
acc. to: Gubanov 1996 |