| Class: | angiosperms |
| Order: | Rosales |
| Family: | Rhamnaceae |
| Genus: | Rhamnus |
| Scientific name: | Rhamnus erythroxylon Pall. |
| Name acc. to: | Gubanov 1996 |
| Herbar: | list records  |
| Description: | Strongly branched spinose shrub up to 2 m high. Leaves linear-lanceolate 5-9 x 0.6-1 cm. Drupe black at maturity, globose, tripartite, 4-6 mm in diameter. |
| Comments: | Plant clearly characterised by small, long leaves. In Flora of China and IPINI as Rhamnus erythroxylum Pallas. |
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| Habitat: | Rocks, stony slopes, granite residual mountains (Grubov 2001). |
| Habit (i)general appearance of a plant | |
| Growth form: (i)Herb, shrub, tree or climber. | shrub, subshrub or semishrub (i)Shrub, multi-stemmed, mostly (0.2) 0.5 - 5 m high, shoots woody up to the tip
example: Caragana leucophloea   inherited by genus Rhamnus: shrub, subshrub or semishrub
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| Size of plant: (i)Attention: use flowering or fruiting specimens to assess plant height (many biennial plants possess only a basal rosette in the first year). | from 1000 mm to 3000 mm inherited by genus Rhamnus:
more than 3000 mm inherited by genus Rhamnus:
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| Parasite status: (i)Is the plant a half- or full parasite? | no parasite/saprophyte (i)Plant fully autonomous, leaves with chlorophyll
example: Most plants, Ranunculus inherited by family Rhamnaceae: no parasite/saprophyte
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| Water or terrestrial plant: (i)Where do the plants grow? | terrestrial (i)Plant grows on dry land
example: Orostachys spinosa inherited by genus Rhamnus: terrestrial
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| Leaf (i)expanded, usually photosynthetic organ of a plant (including phylloclades) | |
| Leaf development: (i)Structure and development of leaves. | with green leaves (i)Plant with green leaves inherited by genus Rhamnus: with green leaves
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| Leaf arrangement: (i)Arrangement of leaves at the stem. | opposite, opposite-decussate (i)Two leaves per node
example: Lamiaceae, e.g. Phlomis   inherited by genus Rhamnus: opposite, opposite-decussate
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| Simple or divided leaves: (i)Are the leaves simple or completely divided in several parts? Blade of the leaf entire or (more or less) deeply dissected. Attention: There are various appearances of the leaf margin (from entire to toothed and lobed). Here, we ignore this and ask only for dissections that separate the leaf for more than one third of its length or width, whatever is smaller. Sometimes, it is difficult to tell apart compound leaves from a shoot system with simple leaves: look for stipulae and/or axillary buds at the ground of the leaves: if only some possess these structures, the others are most likely leaflets of a compound leaf. | simple (i)Non-divided leaf, but margin may be incised nearly to the ground  inherited by genus Rhamnus: simple
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| Shape of blade: (i)Easy for simple leaves. In compound leaves use the general shape of leaflet. Always check the ground for largest leaves of a plant. To be worked out: how to handle pinnate leaves? | elliptic (including ovate and obovate) (i)Elliptic: broadest at the middle and narrower at the two equal ends; ovate: egg-shaped, attached at the broad end; obovate: attached at the narrower end
example: Limosella aquatica  inherited by genus Rhamnus: elliptic (including ovate and obovate)
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| Length of leaves: (i)How long is the leaf, be carefull in compound leaves, measure the complete leaf. | from 21 mm to 50 mm inherited by genus Rhamnus:
more than 50 mm inherited by genus Rhamnus:
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| Leaf apex: (i)Appearance of the tip of leaf resp. leaflets in compound leaves. | acuminate (i)Gradually tapering to a (sharp) point
example: Populus laurifolia?  inherited by genus Rhamnus: acuminate
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| Leaf margin: (i)Structure of leaf margin (or that of a leaflet in case of compound leaves). Attention: Here we ask for the leaf margin, defined as all those dissections that separate the leaf for less than one third of its length or width, whatever is smaller. To be worked out: how to handle margin of pinnate leaves? | serrate / dentate / crenulate (i)Margin saw-like or rounded teethed
example: Betula, Lophanthus (crenulate)  inherited by genus Rhamnus: serrate / dentate / crenulate
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| Petiole: (i)Leaf divided into stalk (petiole) and blade. | with (i)Leaves with petiole (stalk)  inherited by genus Rhamnus: with
shorter than blade (i)Petiol shorter than leaf blade inherited by genus Rhamnus: shorter than blade
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| Stipule: (i)Leaflets at the base of the petiole, these are smaller and of different shape. | pair (i)A pair of free stipulae
example: Lathyrus, Trifolium  inherited by genus Rhamnus: pair
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| Leaf colour upper side: (i)Shades of green on the leaf, upper side. | green (i)Clear green
example: Tribulus terrestris inherited by genus Rhamnus: green
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| Leaf colour lower side: (i)Shades of green on the leaf, lower side. | green (i)Clear green, in most species
example: Angelica decurrens inherited by genus Rhamnus: green
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| Leaf veination: (i)Arrangement of the main veins of a leaf. | pinnate (i)One main vein, several side veins, sometimes inconspicuous
example: Cicerbita    inherited by family Rhamnaceae: pinnate
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| Flower (i)reproductive portion of the plant, consisting of sepals, petals, stamens, and pistils | |
| Flower appearance and pollination: (i)General appearance of the flower. | attractive, animal-pollinated (i)attractive and coloured flowers, mostly large, attracting surely animals
example: Trollius, Rosa, Chamaerhodos inherited by genus Rhamnus: attractive, animal-pollinated
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| Flower colour: (i)Attention: assess colour of the most colourful parts of the flower, but not of the stamens; be aware of single plants with a mutation (mostly white) on flower colour. | greenish (i)petals absent or not distinctly different from colours of leaves, only stigmas (white) or anthers (yellow) may differ in color
example: Chenopodium, Triglochin inherited by genus Rhamnus: greenish
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| Diameter of flower: (i)Diameter of flower or flower head. | to 5 mm (i)
example: Aruncus inherited by genus Rhamnus:
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| Flower symmetry: (i)Symmetry of the perianth leaves. Attention: to assess this character, look on sepals, petals and stamens, but neglect carpels and ovary. | radiary, regular (actinomorphic) (i)More than two axis of symmetry
example: Saxifraga: 5; Iris: 3  inherited by genus Rhamnus: radiary, regular (actinomorphic)
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| Flower form: (i)common forms of flowers ? Veronica | simple (flat) - Do not confuse with inflorescences as in some Asteraceae (i)Petals spread out, flower appearing flat
example: Mollugo, Trientalis, Pulsatilla, Saxifraga  inherited by genus Rhamnus: simple (flat) - Do not confuse with inflorescences as in some Asteraceae
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| Sepal number: (i)Number of sepal leaves (outer perianth leaves, calyx leaves, mostly greenish). Attention, this character applies only for flowers separated in sepals and petals, thus excluding most monocots. Be aware of the bracts (involucral leaves) of Asteraceae flowerheads, do not qualify these as sepals! Be also aware in Rosaceae is often an epicalyx developed, in this case count all parts. | 4 (i)
example: Sinapis inherited by genus Rhamnus: 4
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| Petal / Tepal number: (i)Number of petal leaves (inner perianth leaves, usually coloured). | 4 (i)
example: Galium inherited by genus Rhamnus: 4
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| Petal / Tepal fusion: (i)To which degree are the petal leaves connected? Petals sympetalous. | free (i)all petal leaves separate from each other
example: Anthriscus inherited by genus Rhamnus: free
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| Spur: (i)A hollow, slender, sac-like appendage of the perianth leaves, storing nectar. | no spur (i)Flower without appendage
example: Peganum inherited by family Rhamnaceae: no spur
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| Stamen number: (i)Attention: We ask for the reproductive organs of the flower dispersing pollen. Count only fully fertile stamens, not staminodia (e.g. Parnassia). | 4 (i)Extremely rare, may be absent
example: Plantago inherited by genus Rhamnus: 4
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| Stamen fusion: (i)To which degree are the stamens fused? Attention: Whereas the pollen sacs itself are often free., their stalks (filaments) may be fused. Here, we count them as fused if they are together over at least one thirth of their length. | free (i)Stamens with separate bases
example: Malus inherited by genus Rhamnus: free
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| Pistil number: (i)Number of pistils (female floral organs: style, if developed; stigma and carpels/ovary together build the pistil). | 1 (i)One carpel, but clearly one stigma
example: Pyrola, Primula, Alyssum inherited by genus Rhamnus: 1
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| Carpel fusion: (i)To which degree are the carpels (modified leaf forming simple pistil or part of a compound pistil) fused. | fused (i)Carpels united into an ovary, only styles are free
example: Malus, Berberis inherited by genus Rhamnus: fused
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| Style number: (i)Portion of the pistil connecting the stigma to the ovary. | 1 inherited by genus Rhamnus: 1
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| Ovary position: (i)For entirely or partly fused carpels, describe their position in relation to the insertion point of perianth leaves (best done by doing a longitudinal section of a flower). | superior (hypogynous) (i)Base of carpels attached above insertion point of perianth leaves, carpels free or fused
example: Delphinium, Anemone   inherited by genus Rhamnus: superior (hypogynous)
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| Sex: (i)Distribution of male and female organs among flowers, only most commonly cases. | unisexual (i)
example: Rhodiola inherited by genus Rhamnus: unisexual
dioecious (i)Male and female flowers at different individuals
example: Antennaria inherited by genus Rhamnus: dioecious
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| Inflorescence (i)flowering part of a plant, describes the arrangement of the flowers on the flowering axis | |
| Inflorescence: (i)Structure of the inflorescence. | Flowers in inflorescence (i)No solitary flowers inherited by genus Rhamnus: Flowers in inflorescence
Simple inflorescences (i)Flowers sessile on a main shoot or on short to long not branched side shoots
example: Polygonum bistorta   inherited by genus Rhamnus: Simple inflorescences
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| Appearance: (i)Outer look of the inflorescence. | axillary (i)Usually several inflorescences in axillary shoots or single flowers in leaf axils, main shoot remains mostly leafy
example: Tragopogon, Aconogonon inherited by genus Rhamnus: axillary
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| Inflorescence type: (i)Types of inflorescence. Attention: We here ask for the botanical nomenclature of inflorescences, which is sufficiently complicated. Tick only, if you are certain, or tick all inflorescence types that appear similar of these of the plant in question. | others (in traits_comments nicht aufgeführt) (i)Not as above
example: Sparganium: globose capitate inherited by genus Rhamnus: others (in traits_comments nicht aufgeführt)
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| Fruit (i)the seed bearing organ, with or without adnate parts; a ripened ovary and any other structures which are attached and ripen with it. Aggregate fruits are handled like simple fruits for determination. | |
| Type of fruit: (i)Common fruit types (including pseudocarp). | Indehiscent fruits inherited by genus Rhamnus: Indehiscent fruits
Solitary fruits (i)    inherited by genus Rhamnus: Solitary fruits
drupe (stone fruit) (i)Fleshy, indehiscent fruit with a single, hard stone inside
example: Plum, cherry  inherited by genus Rhamnus: drupe (stone fruit)
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| Size of fruit: (i)Size of the fruit including appendage. | to 5 mm (i)
example: Halerpestes: many folicles forming dry nutlets inherited by genus Rhamnus:
from 5 mm to 10 mm (i)
example: Silene: small capsule opening with teeth inherited by genus Rhamnus:
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| Seed number: (i)Estimate the number of seeds per fruit, if recognizable seeds are in the fruit (in rare cases a fruit may contain one seeded nuts: rose hip, carex). | 2-6 (i)2-6 single seeds, well recognizable
example: Crataegus: few-seeded berry inherited by genus Rhamnus: 2-6
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| Shoot/Stem (i)a young stem or branch | |
| Spines, thorns or prickles: (i)Shoot with conspicuous spines, thorns or prickles. | spines or thornes (i)Sharp pointed woody structure originating from the plant (thornes derived from a reduced branch, spines from leaves)
example: Prunus spinosa no in Mongolia inherited by genus Rhamnus: spines or thornes
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| Root / shoot below ground (i)plant part below ground (in most cases), including below ground shoots, without leaves | |
| Root type: (i)Organisation of the roots. | allorhizous (i)Plant with a conspicuous tap root, one larger tap root with side roots
example: Dicotyledonae inherited by order Rosales: allorhizous
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| Distribution (i)region where the plant is likely to be found | |
| Distribution (Veg. Zones): (i)acc. to Grubov 1952 | Khentei (i)In distribution data often named as '2' 
Khangai (i)In distribution data often named as '3' 
Mongol-Daurian (i)In distribution data often named as '4' 
Middle Khalkha (i)In distribution data often named as '8' 
East Mongolia (i)In distribution data often named as '9' 
East Gobi (i)In distribution data often named as '12' 
Gobi-Altai (i)In distribution data often named as '13' 
acc. to: Gubanov 1996 |
| Distribution Khangay: (i)acc. Flora Khangaya 1989 | I
IV
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