| Class: | angiosperms |
| Order: | Ericales |
| Family: | Ericaceae |
| Genus: | Vaccinium |
| Scientific name: | Vaccinium uliginosum L. |
| Name acc. to: | Gubanov 1996 |
| Herbar: | list records |
| Comments: | See also http://www.manfred-vesper.de/datei.php?did=257 |
| Link to Flora of China: | http://www.efloras.org/browse.aspx?flora_id=2&name_str=Vaccinium+uliginosum |
| open map in a new window |  |
| Habitat: | Goltzy, stone fields, pebbles, dwarf birch and willow thickets, coniferous forests in alpine belt and upper part of forest belt (Grubov 2001). |
| Habit (i)general appearance of a plant | |
| Growth form: (i)Herb, shrub, tree or climber. | shrub, subshrub or semishrub (i)Shrub, multi-stemmed, mostly (0.2) 0.5 - 5 m high, shoots woody up to the tip
example: Caragana leucophloea   inherited by family Ericaceae: shrub, subshrub or semishrub
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| Parasite status: (i)Is the plant a half- or full parasite? | no parasite/saprophyte (i)Plant fully autonomous, leaves with chlorophyll
example: Most plants, Ranunculus inherited by family Ericaceae: no parasite/saprophyte
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| Water or terrestrial plant: (i)Where do the plants grow? | terrestrial (i)Plant grows on dry land
example: Orostachys spinosa inherited by family Ericaceae: terrestrial
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| Leaf (i)expanded, usually photosynthetic organ of a plant (including phylloclades) | |
| Leaf development: (i)Structure and development of leaves. | with green leaves (i)Plant with green leaves inherited by family Ericaceae: with green leaves
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| Leaf arrangement: (i)Arrangement of leaves at the stem. | alternate (i)One leaf per node; distiche: arranged in two vertical rows, equitant
example: Phragmites   inherited by family Ericaceae: alternate
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| Simple or divided leaves: (i)Are the leaves simple or completely divided in several parts? Blade of the leaf entire or (more or less) deeply dissected. Attention: There are various appearances of the leaf margin (from entire to toothed and lobed). Here, we ignore this and ask only for dissections that separate the leaf for more than one third of its length or width, whatever is smaller. Sometimes, it is difficult to tell apart compound leaves from a shoot system with simple leaves: look for stipulae and/or axillary buds at the ground of the leaves: if only some possess these structures, the others are most likely leaflets of a compound leaf. | simple (i)Non-divided leaf, but margin may be incised nearly to the ground  inherited by family Ericaceae: simple
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| Shape of blade: (i)Easy for simple leaves. In compound leaves use the general shape of leaflet. Always check the ground for largest leaves of a plant. To be worked out: how to handle pinnate leaves? | elliptic (including ovate and obovate) (i)Elliptic: broadest at the middle and narrower at the two equal ends; ovate: egg-shaped, attached at the broad end; obovate: attached at the narrower end
example: Limosella aquatica 
round to orbicular (i)Leaf at least as broad as long
example: Pyrola rotundifolia, Caltha 
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| Length of leaves: (i)How long is the leaf, be carefull in compound leaves, measure the complete leaf. | from 6 mm to 10 mm
from 11 mm to 20 mm
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| Width of leaves: (i)How broad is the leaf, be carefull in compound leaves, measure the complete leaf. | 5-10 mm
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| Leaf apex: (i)Appearance of the tip of leaf resp. leaflets in compound leaves. | rounded (i)With a round apex
example: Trifolium 
obtuse (i)Sides coming together at the apex at an angle greater than 90 degrees
example: Fallopia convolvulus 
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| Leaf margin: (i)Structure of leaf margin (or that of a leaflet in case of compound leaves). Attention: Here we ask for the leaf margin, defined as all those dissections that separate the leaf for less than one third of its length or width, whatever is smaller. To be worked out: how to handle margin of pinnate leaves? | entire (i)Plain margin, not toothed
example: Iris 
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| Leaf base: (i)The angle the leaf blade forms with a real or imaginary (sessile leaves) petiole. Attention: Here, we consider only the base of leaves or leaflets (in case of compound leaves). | broad (i)Angle 30° - 90°
example: Cotoneaster 
cuneate (i)Tapering gradually, angle > 90° - 120°
example: Thalictrum, Betula fruticosa 
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| Petiole: (i)Leaf divided into stalk (petiole) and blade. | with (i)Leaves with petiole (stalk) 
shorter than blade (i)Petiol shorter than leaf blade
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| Stipule: (i)Leaflets at the base of the petiole, these are smaller and of different shape. | none (i)Without stipules
example: Euphorbia, Ericaceae s.l. inherited by family Ericaceae: none
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| Leaf colour upper side: (i)Shades of green on the leaf, upper side. | green (i)Clear green
example: Tribulus terrestris inherited by family Ericaceae: green
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| Leaf colour lower side: (i)Shades of green on the leaf, lower side. | green (i)Clear green, in most species
example: Angelica decurrens inherited by genus Vaccinium: green
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| Leaf veination: (i)Arrangement of the main veins of a leaf. | pinnate (i)One main vein, several side veins, sometimes inconspicuous
example: Cicerbita    inherited by family Ericaceae: pinnate
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| Flower (i)reproductive portion of the plant, consisting of sepals, petals, stamens, and pistils | |
| Flower appearance and pollination: (i)General appearance of the flower. | attractive, animal-pollinated (i)attractive and coloured flowers, mostly large, attracting surely animals
example: Trollius, Rosa, Chamaerhodos inherited by family Ericaceae: attractive, animal-pollinated
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| Flower colour: (i)Attention: assess colour of the most colourful parts of the flower, but not of the stamens; be aware of single plants with a mutation (mostly white) on flower colour. | greenish (i)petals absent or not distinctly different from colours of leaves, only stigmas (white) or anthers (yellow) may differ in color
example: Chenopodium, Triglochin
white (i)Most plants of the population white
example: Pleurospermum, Maianthemum
pink (i)Between red and white
example: Centaurium
red (i)Reddish (also orange) to deep red
example: Lilium, Rhododendrum
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| Perianth arrangement: (i)Attention: in some plants, flowers may be dimorphic in different ways (dioecious or gynodioecious). If flowers vary, record the characters of the most showy flowers. | double, different (i)Two types of perianth leaves, differently coloured (sepals: outer periant leaves, usually greenish, and petals: inner perianth leaves, usually coloured)
example: Parnassia   inherited by family Ericaceae: double, different
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| Diameter of flower: (i)Diameter of flower or flower head. | to 5 mm (i)
example: Aruncus
from 5 mm to 10 mm (i)
example: Stellaria
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| Flower symmetry: (i)Symmetry of the perianth leaves. Attention: to assess this character, look on sepals, petals and stamens, but neglect carpels and ovary. | radiary, regular (actinomorphic) (i)More than two axis of symmetry
example: Saxifraga: 5; Iris: 3  inherited by family Ericaceae: radiary, regular (actinomorphic)
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| Sepal number: (i)Number of sepal leaves (outer perianth leaves, calyx leaves, mostly greenish). Attention, this character applies only for flowers separated in sepals and petals, thus excluding most monocots. Be aware of the bracts (involucral leaves) of Asteraceae flowerheads, do not qualify these as sepals! Be also aware in Rosaceae is often an epicalyx developed, in this case count all parts. | 5 (i)
example: Polemonium inherited by family Ericaceae: 5
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| Sepal fusion: (i)To which degree are the sepal leaves connected? Attention, this character applies only for flowers separated in sepals and petals, thus excluding most monocots. Be aware of the bracts (involucral leaves) of Asteraceae flowerheads, do not qualify these as sepals! | free (i)All leaves separate from each other
example: Geranium inherited by family Ericaceae: free
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| Petal / Tepal number: (i)Number of petal leaves (inner perianth leaves, usually coloured). | 5 (i)
example: Potentilla inherited by family Ericaceae: 5
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| Petal / Tepal fusion: (i)To which degree are the petal leaves connected? Petals sympetalous. | fused (i)petal leaves united, only tips are free (gamopetalous, sympetalous)
example: Linnaea, Adenophora, Stellera inherited by family Ericaceae: fused
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| Spur: (i)A hollow, slender, sac-like appendage of the perianth leaves, storing nectar. | no spur (i)Flower without appendage
example: Peganum inherited by family Ericaceae: no spur
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| Stamen number: (i)Attention: We ask for the reproductive organs of the flower dispersing pollen. Count only fully fertile stamens, not staminodia (e.g. Parnassia). | 10 (i)
example: Silene inherited by family Ericaceae: 10
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| Stamen fusion: (i)To which degree are the stamens fused? Attention: Whereas the pollen sacs itself are often free., their stalks (filaments) may be fused. Here, we count them as fused if they are together over at least one thirth of their length. | free (i)Stamens with separate bases
example: Malus inherited by family Ericaceae: free
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| Pistil number: (i)Number of pistils (female floral organs: style, if developed; stigma and carpels/ovary together build the pistil). | 1 (i)One carpel, but clearly one stigma
example: Pyrola, Primula, Alyssum inherited by family Ericaceae: 1
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| Carpel number: (i)Number of carpels (carpel: forming a simple pistil or part of a compound pistil, modified leaf). | 5 inherited by family Ericaceae: 5
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| Carpel fusion: (i)To which degree are the carpels (modified leaf forming simple pistil or part of a compound pistil) fused. | fused (i)Carpels united into an ovary, only styles are free
example: Malus, Berberis inherited by family Ericaceae: fused
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| Style number: (i)Portion of the pistil connecting the stigma to the ovary. | 1 inherited by family Ericaceae: 1
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| Ovary position: (i)For entirely or partly fused carpels, describe their position in relation to the insertion point of perianth leaves (best done by doing a longitudinal section of a flower). | inferior (i)Ovary below the point where perianth leaves are inserted, always fused to an ovary
example: Vaccinum   inherited by genus Vaccinium: inferior
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| Sex: (i)Distribution of male and female organs among flowers, only most commonly cases. | bisexual, hermaphrodite (i)All or nearly all flowers of a plant with male and female parts
example: Haplophyllum, Chenopodium inherited by family Ericaceae: bisexual, hermaphrodite
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| Fruit (i)the seed bearing organ, with or without adnate parts; a ripened ovary and any other structures which are attached and ripen with it. Aggregate fruits are handled like simple fruits for determination. | |
| Consistency: (i)Fleshy fruits or dry fruits, see dispersal adaptations for further classification. | fleshy (i)Usually non dehiscent; with a fleshy (swollen because of a high water content) outer shell, flesh edible at least for animals
example: Prunus, Amygdalus, Malus inherited by genus Vaccinium: fleshy
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| Type of fruit: (i)Common fruit types (including pseudocarp). | Indehiscent fruits inherited by genus Vaccinium: Indehiscent fruits
Solitary fruits (i)    inherited by genus Vaccinium: Solitary fruits
berry (i)Fleshy fruit with several to many seeds in the flesh
example: Tomato  inherited by genus Vaccinium: berry
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| Opening of fruit: (i)Mode of dehiscence at maturity to release seeds. | not opening / indehiscent (i)Fruits remain closed at maturity and disperse with seeds inside
example: Corylus (nut), Vaccinium (berry) inherited by genus Vaccinium: not opening / indehiscent
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| Size of fruit: (i)Size of the fruit including appendage. | from 5 mm to 10 mm (i)
example: Silene: small capsule opening with teeth inherited by genus Vaccinium:
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| Dispersal: (i)Appearance of fruit or seed (if single) and adaptations to dispersal. | Fleshy, edible (i)At least parts with a soft tissue edible at least for some animals (not necessarily for humans) inherited by genus Vaccinium: Fleshy, edible
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| Root / shoot below ground (i)plant part below ground (in most cases), including below ground shoots, without leaves | |
| Root type: (i)Organisation of the roots. | allorhizous (i)Plant with a conspicuous tap root, one larger tap root with side roots
example: Dicotyledonae inherited by order Ericales: allorhizous
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| Distribution (i)region where the plant is likely to be found | |
| Distribution (Veg. Zones): (i)acc. to Grubov 1952 | Khubsgul (i)In distribution data often named as '1' 
Khentei (i)In distribution data often named as '2' 
Khangai (i)In distribution data often named as '3' 
Mongol-Daurian (i)In distribution data often named as '4' 
Khobdo (i)In distribution data often named as '6' 
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| Distribution Khangay: (i)acc. Flora Khangaya 1989 | I
III
IV
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