| Class: | angiosperms |
| Order: | Lamiales |
| Family: | Scrophulariaceae |
| Genus: | Veronica |
| Scientific name: | Veronica pinnata L. |
| Name acc. to: | Gubanov 1996 |
| Herbar: | list records   |
| Synonym: | Pseudolysimachion pinnatum (acc. to (L.) Holub) |
| Comments: | |
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| Habitat: | Steppe and meadow mountain slopes, waterside meadows and pebbles, rocks and stony slopes (Grubov 2001). |
| Habit (i)general appearance of a plant | |
| Growth form: (i)Herb, shrub, tree or climber. | herb (i)Herbaceous, erect plant, up to 2m high, mostly with a leafy shoot; if perennial, shoots die to the ground each season, shoots are not woody
example: Artemisia pectinata   inherited by family Scrophulariaceae: herb
perennial (i)Living for several to many years, as opposed to annual and biennial
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| Size of plant: (i)Attention: use flowering or fruiting specimens to assess plant height (many biennial plants possess only a basal rosette in the first year). | from 100 mm to 250 mm
from 250 mm to 600 mm
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| Parasite status: (i)Is the plant a half- or full parasite? | no parasite/saprophyte (i)Plant fully autonomous, leaves with chlorophyll
example: Most plants, Ranunculus inherited by genus Veronica: no parasite/saprophyte
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| Water or terrestrial plant: (i)Where do the plants grow? | terrestrial (i)Plant grows on dry land
example: Orostachys spinosa inherited by genus Veronica: terrestrial
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| Leaf (i)expanded, usually photosynthetic organ of a plant (including phylloclades) | |
| Leaf development: (i)Structure and development of leaves. | flattened blade (i)Cross-section of lamina flat, plain inherited by genus Veronica: flattened blade
common leaf (i)Green, often divided in blade and petiole
example: Cotoneaster inherited by genus Veronica: common leaf
with green leaves (i)Plant with green leaves inherited by family Scrophulariaceae: with green leaves
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| Leaf arrangement: (i)Arrangement of leaves at the stem. | alternate (i)One leaf per node; distiche: arranged in two vertical rows, equitant
example: Phragmites  
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| Simple or divided leaves: (i)Are the leaves simple or completely divided in several parts? Blade of the leaf entire or (more or less) deeply dissected. Attention: There are various appearances of the leaf margin (from entire to toothed and lobed). Here, we ignore this and ask only for dissections that separate the leaf for more than one third of its length or width, whatever is smaller. Sometimes, it is difficult to tell apart compound leaves from a shoot system with simple leaves: look for stipulae and/or axillary buds at the ground of the leaves: if only some possess these structures, the others are most likely leaflets of a compound leaf. | simple (i)Non-divided leaf, but margin may be incised nearly to the ground  inherited by genus Veronica: simple
pinnatisected, but simple leaf (i)Leaf divided to more than 1/3 of blade; there is a gradual transition from deeply divided leaves to pinnate leaves
example: Papaver 
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| Length of leaves: (i)How long is the leaf, be carefull in compound leaves, measure the complete leaf. | from 11 mm to 20 mm
from 21 mm to 50 mm
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| Width of leaves: (i)How broad is the leaf, be carefull in compound leaves, measure the complete leaf. | 1-2 mm
2-3 mm
3-5 mm
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| Leaf apex: (i)Appearance of the tip of leaf resp. leaflets in compound leaves. | rounded (i)With a round apex
example: Trifolium 
acuminate (i)Gradually tapering to a (sharp) point
example: Populus laurifolia? 
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| Leaf margin: (i)Structure of leaf margin (or that of a leaflet in case of compound leaves). Attention: Here we ask for the leaf margin, defined as all those dissections that separate the leaf for less than one third of its length or width, whatever is smaller. To be worked out: how to handle margin of pinnate leaves? | serrate / dentate / crenulate (i)Margin saw-like or rounded teethed
example: Betula, Lophanthus (crenulate) 
entire (i)Plain margin, not toothed
example: Iris 
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| Leaf base: (i)The angle the leaf blade forms with a real or imaginary (sessile leaves) petiole. Attention: Here, we consider only the base of leaves or leaflets (in case of compound leaves). | narrow (i)Angle < 30° 
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| Petiole: (i)Leaf divided into stalk (petiole) and blade. | with (i)Leaves with petiole (stalk) 
without (i)Leaves without petiole (stalk), sessile
example: Poaceae, Iris 
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| Stipule: (i)Leaflets at the base of the petiole, these are smaller and of different shape. | none (i)Without stipules
example: Euphorbia, Ericaceae s.l. inherited by genus Veronica: none
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| Leaf colour upper side: (i)Shades of green on the leaf, upper side. | green (i)Clear green
example: Tribulus terrestris
grayish (i)Grayish in colour due to hairs or a thick cuticula
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| Leaf colour lower side: (i)Shades of green on the leaf, lower side. | green (i)Clear green, in most species
example: Angelica decurrens
grayish (i)Grayish in colour due to hairs or a thick cuticula
example: Atriplex cana
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| Leaf veination: (i)Arrangement of the main veins of a leaf. | pinnate (i)One main vein, several side veins, sometimes inconspicuous
example: Cicerbita    inherited by genus Veronica: pinnate
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| Flower (i)reproductive portion of the plant, consisting of sepals, petals, stamens, and pistils | |
| Flower appearance and pollination: (i)General appearance of the flower. | attractive, animal-pollinated (i)attractive and coloured flowers, mostly large, attracting surely animals
example: Trollius, Rosa, Chamaerhodos inherited by family Scrophulariaceae: attractive, animal-pollinated
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| Flower colour: (i)Attention: assess colour of the most colourful parts of the flower, but not of the stamens; be aware of single plants with a mutation (mostly white) on flower colour. | white (i)Most plants of the population white
example: Pleurospermum, Maianthemum inherited by genus Veronica: white
pink (i)Between red and white
example: Centaurium inherited by genus Veronica: pink
blue (i)Bluish colours with mixture to red to deep blue
example: Myosotis, Dracocephalum inherited by genus Veronica: blue
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| Perianth arrangement: (i)Attention: in some plants, flowers may be dimorphic in different ways (dioecious or gynodioecious). If flowers vary, record the characters of the most showy flowers. | double, different (i)Two types of perianth leaves, differently coloured (sepals: outer periant leaves, usually greenish, and petals: inner perianth leaves, usually coloured)
example: Parnassia   inherited by family Scrophulariaceae: double, different
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| Diameter of flower: (i)Diameter of flower or flower head. | from 5 mm to 10 mm (i)
example: Stellaria
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| Flower symmetry: (i)Symmetry of the perianth leaves. Attention: to assess this character, look on sepals, petals and stamens, but neglect carpels and ovary. | zygomorphic (i)One axis of symmetry, monosymmetrical flowers
example: Pedicularis, Nepeta, Viola    inherited by genus Veronica: zygomorphic
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| Flower form: (i)common forms of flowers ? Veronica | simple (flat) - Do not confuse with inflorescences as in some Asteraceae (i)Petals spread out, flower appearing flat
example: Mollugo, Trientalis, Pulsatilla, Saxifraga  inherited by genus Veronica: simple (flat) - Do not confuse with inflorescences as in some Asteraceae
tubular to funnel-shaped (i)Petals form a tube, are often partially united to a cylindrical corolla, often surrounded by a calyx  inherited by genus Veronica: tubular to funnel-shaped
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| Sepal number: (i)Number of sepal leaves (outer perianth leaves, calyx leaves, mostly greenish). Attention, this character applies only for flowers separated in sepals and petals, thus excluding most monocots. Be aware of the bracts (involucral leaves) of Asteraceae flowerheads, do not qualify these as sepals! Be also aware in Rosaceae is often an epicalyx developed, in this case count all parts. | 4 (i)
example: Sinapis inherited by genus Veronica: 4
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| Sepal fusion: (i)To which degree are the sepal leaves connected? Attention, this character applies only for flowers separated in sepals and petals, thus excluding most monocots. Be aware of the bracts (involucral leaves) of Asteraceae flowerheads, do not qualify these as sepals! | fused (i)Leaves united, only tips are free
example: Fabaceae, Silene   inherited by genus Veronica: fused
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| Petal / Tepal number: (i)Number of petal leaves (inner perianth leaves, usually coloured). | 4 (i)
example: Galium inherited by genus Veronica: 4
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| Petal / Tepal fusion: (i)To which degree are the petal leaves connected? Petals sympetalous. | fused (i)petal leaves united, only tips are free (gamopetalous, sympetalous)
example: Linnaea, Adenophora, Stellera inherited by family Scrophulariaceae: fused
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| Spur: (i)A hollow, slender, sac-like appendage of the perianth leaves, storing nectar. | no spur (i)Flower without appendage
example: Peganum inherited by genus Veronica: no spur
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| Stamen number: (i)Attention: We ask for the reproductive organs of the flower dispersing pollen. Count only fully fertile stamens, not staminodia (e.g. Parnassia). | 2 (i)
example: Cypripedium inherited by genus Veronica: 2
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| Stamen fusion: (i)To which degree are the stamens fused? Attention: Whereas the pollen sacs itself are often free., their stalks (filaments) may be fused. Here, we count them as fused if they are together over at least one thirth of their length. | fused with a corolla (calyx in Thymelaeaceae) (i)Stamens with perianth leaves at least one third of the length of the filament
example: Orobanche, Salvia, Stellera inherited by family Scrophulariaceae: fused with a corolla (calyx in Thymelaeaceae)
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| Pistil number: (i)Number of pistils (female floral organs: style, if developed; stigma and carpels/ovary together build the pistil). | 1 (i)One carpel, but clearly one stigma
example: Pyrola, Primula, Alyssum inherited by genus Veronica: 1
2 (i)Two stigmas, often cleaved like a snakes tongue
example: Salvia, Arnica, Bupleurum, Bromus, Saxifraga, Veronica inherited by genus Veronica: 2
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| Carpel number: (i)Number of carpels (carpel: forming a simple pistil or part of a compound pistil, modified leaf). | 2 inherited by family Scrophulariaceae: 2
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| Carpel fusion: (i)To which degree are the carpels (modified leaf forming simple pistil or part of a compound pistil) fused. | fused (i)Carpels united into an ovary, only styles are free
example: Malus, Berberis inherited by genus Veronica: fused
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| Style number: (i)Portion of the pistil connecting the stigma to the ovary. | 1 inherited by family Scrophulariaceae: 1
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| Ovary position: (i)For entirely or partly fused carpels, describe their position in relation to the insertion point of perianth leaves (best done by doing a longitudinal section of a flower). | superior (hypogynous) (i)Base of carpels attached above insertion point of perianth leaves, carpels free or fused
example: Delphinium, Anemone   inherited by family Scrophulariaceae: superior (hypogynous)
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| Sex: (i)Distribution of male and female organs among flowers, only most commonly cases. | bisexual, hermaphrodite (i)All or nearly all flowers of a plant with male and female parts
example: Haplophyllum, Chenopodium inherited by family Scrophulariaceae: bisexual, hermaphrodite
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| Inflorescence (i)flowering part of a plant, describes the arrangement of the flowers on the flowering axis | |
| Inflorescence: (i)Structure of the inflorescence. | Flowers in inflorescence (i)No solitary flowers inherited by genus Veronica: Flowers in inflorescence
Simple inflorescences (i)Flowers sessile on a main shoot or on short to long not branched side shoots
example: Polygonum bistorta  
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| Appearance: (i)Outer look of the inflorescence. | terminal (i)Inflorescence is the highest point of the plant and may consist of a single flower only
example: Cypripedium, Rhaponticum, Ligularia sibirica, Echinops
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| Inflorescence type: (i)Types of inflorescence. Attention: We here ask for the botanical nomenclature of inflorescences, which is sufficiently complicated. Tick only, if you are certain, or tick all inflorescence types that appear similar of these of the plant in question. | spike (spadix) (i)All flowers sessile and crowded along a main axis, there may be several spikes on a shoot; sometimes axis thickened (spadix)
example: Plantago, Carex vesicaria, Vicia, Typha (spadix)  
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| Fruit (i)the seed bearing organ, with or without adnate parts; a ripened ovary and any other structures which are attached and ripen with it. Aggregate fruits are handled like simple fruits for determination. | |
| Consistency: (i)Fleshy fruits or dry fruits, see dispersal adaptations for further classification. | dry (i)With a dry outer shell, no fleshy parts, but seed (embryo) could be edible inherited by family Scrophulariaceae: dry
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| Type of fruit: (i)Common fruit types (including pseudocarp). | Solitary fruits (i)    inherited by family Scrophulariaceae: Solitary fruits
capsule (i)Dry dehiscent fruit, releasing seeds by slits or holes.
example: Poppy, most Caryophyllaceae, Cerastium, a lot of Scrophulariaceae, Iris (oppened capsule looks like Delphinium), Zygophyllum - it is a very common fruit type    inherited by family Scrophulariaceae: capsule
Dehiscent fruits (i)Fruits open along a longitudinale line (except silicula) inherited by family Scrophulariaceae: Dehiscent fruits
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| Opening of fruit: (i)Mode of dehiscence at maturity to release seeds. | opening along dehiscent line (i)Opening along a preformed line
example: Vicia, Lathyrus: pods inherited by genus Veronica: opening along dehiscent line
opening / dehiscent (i)Dry? Fruits opening with different types inherited by genus Veronica: opening / dehiscent
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| Size of fruit: (i)Size of the fruit including appendage. | to 5 mm (i)
example: Halerpestes: many folicles forming dry nutlets
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| Dispersal: (i)Appearance of fruit or seed (if single) and adaptations to dispersal. | Otherwise (i)All parts dry, no conspicuous adaptations
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| Seed number: (i)Estimate the number of seeds per fruit, if recognizable seeds are in the fruit (in rare cases a fruit may contain one seeded nuts: rose hip, carex). | 7-50 (i)Numerable, but may be counted
example: Vaccinum: multi-seeded berry, Ptilotrichum: few-seeded siliqua inherited by genus Veronica: 7-50
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| Hairs | |
| Has hairs?: | has hairs
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| Hairs: (i)Appearance, structure, coverage of hairs on plant. | on leaf (i)Hairs on upper side, lower side or on margin of leaf
coverage: scattered (i)Solitary hairs in large distances to each other
example: Dracocephalum origanoides
leaf upper side (i)Has hairs on leaves upper side (blade)
coverage: scattered (i)Solitary hairs in large distances to each other
example: Arabis
appearance: soft (i)Hairs very flexible and soft, lay down at a touch
example: Lonicera xylosteum
appearance: soft (i)Hairs very flexible and soft, lay down at a touch
example: Rubus
leaf lower side
on stem/shoot (i)Has hairs on stem/shoot
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| Shoot/Stem (i)a young stem or branch | |
| Spines, thorns or prickles: (i)Shoot with conspicuous spines, thorns or prickles. | absent (i)Stem glabrous or hairy, but never with spines, thornes or prickles
example: Gentiana barbata inherited by genus Veronica: absent
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| Root / shoot below ground (i)plant part below ground (in most cases), including below ground shoots, without leaves | |
| Root type: (i)Organisation of the roots. | allorhizous (i)Plant with a conspicuous tap root, one larger tap root with side roots
example: Dicotyledonae inherited by order Lamiales: allorhizous
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| Runners: (i)Plant must be excavated; shoots, subterranean shoots connected by runners. | long root suckers or rhizomes (i)Plants grow new shoots from roots or subterranean shoots, called rhizomes, these spacing stems apart
example: Hippophae, Artemisia sericea
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| Storage in below-ground structures: (i)Rhizomes or bulbs. | none (i)Plant with non-thickened roots, their maximum diameter does not exceed the diameter of the shoot base
example: Anabasis brevifolia inherited by genus Veronica: none
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| Distribution (i)region where the plant is likely to be found | |
| Distribution (Veg. Zones): (i)acc. to Grubov 1952 | Khentei (i)In distribution data often named as '2' 
Khangai (i)In distribution data often named as '3' 
Mongol-Daurian (i)In distribution data often named as '4' 
Khobdo (i)In distribution data often named as '6' 
Mongolian Altai (i)In distribution data often named as '7' 
Depression of Great Lakes (i)In distribution data often named as '10' 
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| Distribution Khangay: (i)acc. Flora Khangaya 1989 | I
II
III
V
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