| Class: | angiosperms |
| Order: | Zygophyllales |
| Family: | Zygophyllaceae |
| Genus: | Tribulus |
| Scientific name: | Tribulus terrestris L. |
| Name acc. to: | Gubanov 1996 |
| Herbar: | list records    |
| Link to Flora of China: | http://www.efloras.org/browse.aspx?flora_id=2&name_str=Tribulus+terrestris |
| open map in a new window |  |
| Habitat: | Alkaline, debris and stony tailings, sands, sandy bottom of sayrs, alkaline river and spring banks, toyrims, ruderal places, banks of irrigation ditches, along roads and paths (Grubov 2001). |
| Habit (i)general appearance of a plant | |
| Growth form: (i)Herb, shrub, tree or climber. | annual (i)Completing its life cycle within one year or one growing season; roots weak and thin
herb (i)Herbaceous, erect plant, up to 2m high, mostly with a leafy shoot; if perennial, shoots die to the ground each season, shoots are not woody
example: Artemisia pectinata 
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| Size of plant: (i)Attention: use flowering or fruiting specimens to assess plant height (many biennial plants possess only a basal rosette in the first year). | from 100 mm to 250 mm
from 250 mm to 600 mm
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| Parasite status: (i)Is the plant a half- or full parasite? | no parasite/saprophyte (i)Plant fully autonomous, leaves with chlorophyll
example: Most plants, Ranunculus  inherited by family Zygophyllaceae: no parasite/saprophyte
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| Water or terrestrial plant: (i)Where do the plants grow? | terrestrial (i)Plant grows on dry land
example: Orostachys spinosa inherited by family Zygophyllaceae: terrestrial
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| Leaf (i)expanded, usually photosynthetic organ of a plant (including phylloclades) | |
| Leaf development: (i)Structure and development of leaves. | with green leaves (i)Plant with green leaves
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| Leaf arrangement: (i)Arrangement of leaves at the stem. | alternate (i)One leaf per node; distiche: arranged in two vertical rows, equitant
example: Phragmites  
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| Simple or divided leaves: (i)Are the leaves simple or completely divided in several parts? Blade of the leaf entire or (more or less) deeply dissected. Attention: There are various appearances of the leaf margin (from entire to toothed and lobed). Here, we ignore this and ask only for dissections that separate the leaf for more than one third of its length or width, whatever is smaller. Sometimes, it is difficult to tell apart compound leaves from a shoot system with simple leaves: look for stipulae and/or axillary buds at the ground of the leaves: if only some possess these structures, the others are most likely leaflets of a compound leaf. | paripinnate / evenpinnate (i)A pinnate leaf with all leaflets in pairs
compound (i)Composed of several similar parts
pinnate (i)With leaflets on both sides of an axis, at least 2 pairs or more
example: Onobrychis 
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| Length of leaves: (i)How long is the leaf, be carefull in compound leaves, measure the complete leaf. | from 21 mm to 50 mm
more than 50 mm
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| Leaf margin: (i)Structure of leaf margin (or that of a leaflet in case of compound leaves). Attention: Here we ask for the leaf margin, defined as all those dissections that separate the leaf for less than one third of its length or width, whatever is smaller. To be worked out: how to handle margin of pinnate leaves? | entire (i)Plain margin, not toothed
example: Iris 
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| Petiole: (i)Leaf divided into stalk (petiole) and blade. | with (i)Leaves with petiole (stalk) 
shorter than blade (i)Petiol shorter than leaf blade
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| Stipule: (i)Leaflets at the base of the petiole, these are smaller and of different shape. | pair (i)A pair of free stipulae
example: Lathyrus, Trifolium 
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| Leaf colour upper side: (i)Shades of green on the leaf, upper side. | green (i)Clear green
example: Tribulus terrestris inherited by genus Tribulus: green
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| Leaf veination: (i)Arrangement of the main veins of a leaf. | pinnate (i)One main vein, several side veins, sometimes inconspicuous
example: Cicerbita    inherited by family Zygophyllaceae: pinnate
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| Flower (i)reproductive portion of the plant, consisting of sepals, petals, stamens, and pistils | |
| Flower appearance and pollination: (i)General appearance of the flower. | attractive, animal-pollinated (i)attractive and coloured flowers, mostly large, attracting surely animals
example: Trollius, Rosa, Chamaerhodos
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| Flower colour: (i)Attention: assess colour of the most colourful parts of the flower, but not of the stamens; be aware of single plants with a mutation (mostly white) on flower colour. | yellow to orange (i)Pale to golden yellow
example: Ranunculus, Crepis
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| Perianth arrangement: (i)Attention: in some plants, flowers may be dimorphic in different ways (dioecious or gynodioecious). If flowers vary, record the characters of the most showy flowers. | double, different (i)Two types of perianth leaves, differently coloured (sepals: outer periant leaves, usually greenish, and petals: inner perianth leaves, usually coloured)
example: Parnassia  
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| Diameter of flower: (i)Diameter of flower or flower head. | from 5 mm to 10 mm (i)
example: Stellaria
from 10 mm to 20 mm (i)
example: Potentilla
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| Flower symmetry: (i)Symmetry of the perianth leaves. Attention: to assess this character, look on sepals, petals and stamens, but neglect carpels and ovary. | radiary, regular (actinomorphic) (i)More than two axis of symmetry
example: Saxifraga: 5; Iris: 3 
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| Flower form: (i)common forms of flowers ? Veronica | simple (flat) - Do not confuse with inflorescences as in some Asteraceae (i)Petals spread out, flower appearing flat
example: Mollugo, Trientalis, Pulsatilla, Saxifraga 
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| Sepal number: (i)Number of sepal leaves (outer perianth leaves, calyx leaves, mostly greenish). Attention, this character applies only for flowers separated in sepals and petals, thus excluding most monocots. Be aware of the bracts (involucral leaves) of Asteraceae flowerheads, do not qualify these as sepals! Be also aware in Rosaceae is often an epicalyx developed, in this case count all parts. | 5 (i)
example: Polemonium
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| Petal / Tepal number: (i)Number of petal leaves (inner perianth leaves, usually coloured). | 5 (i)
example: Potentilla
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| Petal / Tepal fusion: (i)To which degree are the petal leaves connected? Petals sympetalous. | free (i)all petal leaves separate from each other
example: Anthriscus
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| Spur: (i)A hollow, slender, sac-like appendage of the perianth leaves, storing nectar. | no spur (i)Flower without appendage
example: Peganum inherited by family Zygophyllaceae: no spur
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| Stamen number: (i)Attention: We ask for the reproductive organs of the flower dispersing pollen. Count only fully fertile stamens, not staminodia (e.g. Parnassia). | 10 (i)
example: Silene
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| Stamen fusion: (i)To which degree are the stamens fused? Attention: Whereas the pollen sacs itself are often free., their stalks (filaments) may be fused. Here, we count them as fused if they are together over at least one thirth of their length. | free (i)Stamens with separate bases
example: Malus
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| Pistil number: (i)Number of pistils (female floral organs: style, if developed; stigma and carpels/ovary together build the pistil). | 1 (i)One carpel, but clearly one stigma
example: Pyrola, Primula, Alyssum
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| Carpel number: (i)Number of carpels (carpel: forming a simple pistil or part of a compound pistil, modified leaf). | 5
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| Carpel fusion: (i)To which degree are the carpels (modified leaf forming simple pistil or part of a compound pistil) fused. | fused (i)Carpels united into an ovary, only styles are free
example: Malus, Berberis
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| Style number: (i)Portion of the pistil connecting the stigma to the ovary. | 1
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| Ovary position: (i)For entirely or partly fused carpels, describe their position in relation to the insertion point of perianth leaves (best done by doing a longitudinal section of a flower). | superior (hypogynous) (i)Base of carpels attached above insertion point of perianth leaves, carpels free or fused
example: Delphinium, Anemone  
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| Sex: (i)Distribution of male and female organs among flowers, only most commonly cases. | bisexual, hermaphrodite (i)All or nearly all flowers of a plant with male and female parts
example: Haplophyllum, Chenopodium
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| Inflorescence (i)flowering part of a plant, describes the arrangement of the flowers on the flowering axis | |
| Inflorescence: (i)Structure of the inflorescence. | Solitary flowers (i)Each flower grows on an own leafy stem there may be more than one, if the plant has many leafy shoots
example: Viola, Saxifraga hirculus, Rubus arcticus
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| Appearance: (i)Outer look of the inflorescence. | axillary (i)Usually several inflorescences in axillary shoots or single flowers in leaf axils, main shoot remains mostly leafy
example: Tragopogon, Aconogonon
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| Fruit (i)the seed bearing organ, with or without adnate parts; a ripened ovary and any other structures which are attached and ripen with it. Aggregate fruits are handled like simple fruits for determination. | |
| Consistency: (i)Fleshy fruits or dry fruits, see dispersal adaptations for further classification. | dry (i)With a dry outer shell, no fleshy parts, but seed (embryo) could be edible
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| Type of fruit: (i)Common fruit types (including pseudocarp). | Indehiscent fruits
Solitary fruits (i)   
capsule (i)Dry dehiscent fruit, releasing seeds by slits or holes.
example: Poppy, most Caryophyllaceae, Cerastium, a lot of Scrophulariaceae, Iris (oppened capsule looks like Delphinium), Zygophyllum - it is a very common fruit type   
nut or nutlet (i)Dry fruit with a single, hard stone inside (and usually a large often edible embryo) 
Dehiscent fruits (i)Fruits open along a longitudinale line (except silicula)
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| Opening of fruit: (i)Mode of dehiscence at maturity to release seeds. | not opening / indehiscent (i)Fruits remain closed at maturity and disperse with seeds inside
example: Corylus (nut), Vaccinium (berry)
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| Splitting of fruit: (i)Is a solitary fruit splitting into fruitlets? The fruit splits into separate one-seeded segments at maturity (schizocarp). | splitting
in 4 parts (i)
example: Boraginaceae, Lamiaceae 
in 5 parts (i)
example: Geraniaceae 
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| Size of fruit: (i)Size of the fruit including appendage. | from 5 mm to 10 mm (i)
example: Silene: small capsule opening with teeth
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| Dispersal: (i)Appearance of fruit or seed (if single) and adaptations to dispersal. | Bur (i)Fruit with prickly appendages sticking to fur of animals
example: Lappula, Agrimonia, Bidens
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| Seed number: (i)Estimate the number of seeds per fruit, if recognizable seeds are in the fruit (in rare cases a fruit may contain one seeded nuts: rose hip, carex). | 1 (i)A single seed (stone) or seed and fruit wall tightly connected
example: Prunus, Amygdalus: drupe
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| Hairs | |
| Has hairs?: | has hairs
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| Hairs: (i)Appearance, structure, coverage of hairs on plant. | on leaf (i)Hairs on upper side, lower side or on margin of leaf
leaf upper side (i)Has hairs on leaves upper side (blade)
appearance spiny (i)With spines
example: Tribulus inherited by genus Tribulus: appearance spiny
Hairs on fruits inherited by genus Tribulus: Hairs on fruits
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| Root / shoot below ground (i)plant part below ground (in most cases), including below ground shoots, without leaves | |
| Root type: (i)Organisation of the roots. | allorhizous (i)Plant with a conspicuous tap root, one larger tap root with side roots
example: Dicotyledonae inherited by order Zygophyllales: allorhizous
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| Distribution (i)region where the plant is likely to be found | |
| Distribution (Veg. Zones): (i)acc. to Grubov 1952 | Khangai (i)In distribution data often named as '3' 
Great Khingan (i)In distribution data often named as '5' 
Khobdo (i)In distribution data often named as '6' 
Mongolian Altai (i)In distribution data often named as '7' 
Middle Khalkha (i)In distribution data often named as '8' 
East Mongolia (i)In distribution data often named as '9' 
Depression of Great Lakes (i)In distribution data often named as '10' 
Valley of Lakes (i)In distribution data often named as '11' 
East Gobi (i)In distribution data often named as '12' 
Gobi-Altai (i)In distribution data often named as '13' 
Dzungarian Gobi (i)In distribution data often named as '14' 
Transaltai Gobi (i)In distribution data often named as '15' 
Alashan Gobi (i)In distribution data often named as '16' 
acc. to: Gubanov 1996 |
| Distribution Khangay: (i)acc. Flora Khangaya 1989 | II
VI
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| Plant Use | |
| General Use: | medical plant
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